REFLECTIONS ON THE PRECEDING EXPERIMENTS.
By the experiments we have just described, I was assured that females,
as well as males, have a seminal liquor which contains moving
substances; that these substances were not real animals, but only
living organic particles; and that those particles exist, not only in
the seminal liquors of the two sexes, but even in the flesh of animals,
and in the germs of vegetables. To discover whether all the parts of
animals, and all the germs of vegetables, contained living organic
particles, I caused infusions of the flesh of different animals to
be made, and of more than twenty kinds of seeds of different plants;
and after they had infused four or five days, in phials closely stopt
up, I had the satisfaction to see moving organic parts in them all;
some appeared sooner, and others later; some preserved their motion
for months together, while others were soon deprived of it; some
directly produced large moving globules, that had the appearance of
real animals, which changed their figures, separated, and became
successively smaller: others produced only small globules, whose
motions were very brisk; others produced filaments which lengthened
and seemed to vegetate, swelled, and afterwards thousands of moving
globules issued therefrom; but it is useless to detail my observations
on the infusion of plants, since Mr. Needham has published so excellent
a treatise on the subject. I read the preceding treatise to that able
naturalist, and often reasoned with him on the subject, particularly
on the probability that the germs of vegetables contained similar
moving bodies to those in the seed of male and female animals. He
thought those views sufficiently founded to deserve to be pursued; and
therefore began to make experiments on all parts of vegetables; and I
must own that the ideas I gave him on this subject have reaped greater
profit under his hands than they would have done from me. I could quote
many examples, but shall confine myself to one, because I indicated the
circumstance I am going to relate.
To determine whether the moving substances seen in the infusions
of flesh were true animals, or only, as I supposed, moving organic
particles, Mr. Needham imagined that he had only to examine some
roasted meat, because if they were animals the fire must destroy
them; and if not animals, they might still be found there as well as
when the meat was raw; having therefore taken the jelly of veal, and
other roasted meat, he infused them for several days in water, closely
corked up in phials, and upon examination he found in every one of
them a great quantity of moving substances. He shewed me some of these
infusions, and among the rest that of the jelly of veal, in which there
were moving substances, perfectly like those in the seminal liquor of
a man, a dog, and a bitch, when they have no threads, or tails; and
although we perceived them to change their figures, their motions so
perfectly resembled those of an animal which swims, that whoever saw
them, without being acquainted with what has been already mentioned,
might certainly have taken them for real animals. I shall only add,
that Mr. Needham assured himself, by a multiplicity of experiments,
that all parts of vegetables contain moving organic particles, which
confirms what I have said, and extends my theory on the composition of
organized beings, and their reproduction.
All animals, both male and female, and all vegetables whatsoever, it is
therefore evident are composed of living organic parts. These organic
parts are in the greatest abundance in the seminal liquor of animals,
and in seeds of vegetables. It is from the union of these organic
parts returned from all parts of the animal or vegetable body, that
reproduction is performed, and is always like the animal or vegetable
in which it operates; because the union of these organic parts cannot
be made but by the means of an internal mould, in which the form of
an animal or vegetable is produced. It is in this also the essence of
the unity and continuity of the species consists, and will so continue
while the great Creator permits their existence.
But before I draw general conclusions from the system I am
establishing, I must endeavour to remove some objections which might be
made, and mention some other circumstances which will serve to place
this matter in a better light.
It will be asked, why I deny those moving substances in the seminal
liquors to be animals, since they have constantly been regarded as
such by Leeuwenhoek, and every other naturalist, who has examined
them? I may also be told, that living organic particles are not
perfectly intelligible, if they are to be looked upon as animalculæ;
and to suppose an animal is composed of a number of small animals,
is nearly the same as saying that an organized being is composed of
living organic particles. I shall therefore endeavour to answer these
objections in a satisfactory manner.
It is certain that almost all naturalists agree in looking on the
moving substances in seminal liquors as real animals; but it is no less
certain, from my own observations, and those of Mr. Needham, on the
seed of the calmar, that these moving substances are more simple and
less organized beings than animals.
The word _animal_, in the acceptation we commonly receive it,
represents a general idea formed of particular ideas drawn from
particular animals. All general ideas include many different ones,
which approach, or are more or less distant from each other, and
consequently no general idea can either be exact or precise. The
general idea which we form of an animal may be taken principally from
the particular idea of a dog, a horse, and other beasts, which appear
to us to act and move according to the impulse of their will, and which
are besides composed of flesh and blood, seek after their food, have
sexes, and the faculty of reproduction. The general idea, therefore,
expressed by the word _animal_, must comprehend a number of particular
ideas, not one of which constitutes the essence of the general idea,
for there are animals which appear to have no reason, will, progressive
motion, flesh nor blood, and which only appear to be a congealed
substance: there are some which cannot seek their food, but only
receive it from the element they live in: there are some which have no
sensation, not even that of feeling, at least in any sensible degree:
there are some have no sexes, or are both in one; there only belongs,
therefore, to the animal a general idea of what is common also to the
vegetable, that is, the faculty of reproduction.
The general idea then is formed from the whole taken together, which
whole being composed of different parts, there is consequently between
these parts degrees and links. An insect, in this sense, is something
less of an animal than a dog; an oyster still less than an insect; a
sea-nettle, or a fresh-water polypus, still less than an oyster; and
as nature acts by insensible links, we may find beings which are still
less animated than a sea-nettle, or a polypus. Our general ideas are
only artificial methods to collect a quantity of objects in the same
point of view; and they have, like the artificial methods we shall
speak of, the defect of never being able to comprehend the whole.
They are likewise opposite to the walk of nature, which is uniform,
insensible, and always particular, insomuch that by our endeavouring to
comprehend too great a number of particular ideas in one single word,
we have no longer a clear idea of what that word conveys; because, the
word being received, we imagine that it is a line drawn between the
productions of nature; that all above this line is _animal_, and all
below it _vegetable_; another word, as general as the first, and which
is used as a line of separation between organized bodies and inanimate
matter. But as we have already said, these lines of separation do
not exist in nature; there are beings which are neither animals,
vegetables, nor minerals, and which we in vain might attempt to arrange
with either. For example, when Mr. Trembly first observed the polypus,
he employed a considerable time before he could determine whether it
was an animal or a plant; and possibly from this reason that it is
perhaps neither one nor the other, and all that can be said is, that it
approaches nearest to an animal; and as we suppose every living thing
must be either an animal or a plant, we do not credit the existence of
an organized being, that cannot be referred to one of those general
names; whereas there must, and in fact are, a great number of organized
beings which are neither the one nor the other. The moving substances
perceived in seminal liquors, in infusions of the flesh of animals, in
seed, and other parts of plants, are all of this kind. We cannot call
these animals, nor can we say they are vegetables, and certainly we can
still less assert they are minerals.
We can therefore affirm, without fear of advancing too much, that the
grand division of nature's productions into _Animals_, _Vegetables_,
and _Minerals_, do not contain every material being; since there are
some that exist which cannot be classed in this division. We have
already observed, that nature passes by insensible links from the
animal to the vegetable, but from the vegetable to the mineral the
passage is quick, and the distance considerable; from whence the law of
nature's passing by imperceptible degrees appears untrue. This made me
suppose that by examining nature closely we shall discover intermediate
organized beings, which without having the power of reproduction, like
animals and vegetables, would nevertheless have a kind of life and
motion; other beings which, without being either vegetables or animals,
might possibly enter into the composition of both, and likewise other
beings which would be only the assemblage of the organic molecules I
have spoken of in the preceding chapters.
In the first class of these kind of beings eggs must be placed; those
of hens, and other birds, are fastened to a common pedicle, and draw
their nourishment and growth from the body of the animal, but when
fastened to the ovary, they are not then real eggs, but only yellow
globules which separate from the ovary as soon as they have attained a
certain growth. Their internal organization is such that they derive
nourishment from the lymph, the matrix of the hen, and by which they
form the white membranes, and at last the shell. The egg therefore has
a kind of life and organization, a growth, expansion, and a form which
it assumes by its own powers. It does not live like an animal, nor
vegetate like a plant, nor is possessed of the power of reproduction;
nevertheless it grows, acts externally, and is organized. Must we not
then look upon it as a being of a separate class, and which ought not
to be ranked either with animal or mineral? for if it is pretended that
the egg is only an animal production, destined for the nutriment of
the chicken, and should be looked upon as a part of the hen; I answer,
that the eggs, whether impregnated or not, will be always organized
after the same mode: that impregnation only changes an almost invisible
part; and that it attains its perfection and growth, as well externally
as internally, whether it contains the chicken or not, and that
consequently it ought to be considered as a separate being.
What I have said will appear more clear, if we consider the formation
and growth of the eggs of fish; when the female deposits them in the
water they are only the outlines of eggs, which being separated from
the body of the animal, attract and appropriate to themselves the
particles which agree the best for their nourishment, and grow thus
by intussusception. In the same manner as the hen's egg acquires the
white and membranes in the matrix, wherein it floats, so the eggs of
fish acquire their membranes and white in the water; and whether the
male impregnates them, by emitting on these the liquor of its roe, or
whether they remain unimpregnated, they do not the less attain their
entire perfection. It appears to me, therefore, that the eggs should
be considered as organized bodies, which being neither animals nor
vegetables, are a genus apart.
A second class of beings, of the same kind, are the organized bodies
found in the semen of all animals, and which, like those in the milt of
a calmar, are rather natural machines than animals. These are properly
the first assemblages which result from the organic molecules we have
so much spoken of, and they are, perhaps, the parts which constitute
the organized bodies of animals. They are found in the semen of all
animals, because the semen is only the residue of the organic molecules
that the animal takes in with its aliment, and which, as we have
already observed, are those parts most analogous to the animal itself,
and most organic; it is those particles which compose the matter of the
semen, and consequently we must not be astonished to find organized
bodies therein.
To be perfectly convinced that these organized bodies are not real
animals, we need only reflect on the preceding experiments. The moving
bodies in the seminal liquor have been taken for animals, because they
have a progressive motion, and are thought to have a tail; but if we
consider, on one hand, the nature of this progressive motion, which
finishes in a very short time without ever renewing its motion; and
on the other, the nature of these tails, which are only threads which
the moving bodies draw after them, we shall begin to hesitate; for an
animal goes sometimes slow, sometimes fast, and sometimes remains in
a state of rest; these moving bodies, on the contrary, always continue
the same motion, and I have never seen them stop and renew their
movement again. I ask, whether this kind of continued motion, without
any rest, is common to animals, and if that ought not to make us doubt
these moving bodies being real animals? An animal of any kind must also
have a constant form and distinct limbs; but these moving bodies vary,
and change their forms every moment, have no distinct limbs, and their
tails appear as a part which does not belong to the individual. Can
we then imagine these bodies to be real animals? In seminal liquors
filaments are seen which lengthen and appear to vegetate; after which
they swell and produce moving bodies. These filaments may be kinds
of vegetables, but the moving bodies which spring from them cannot
be animals, for a vegetable has never yet been seen to produce an
animal. These moving bodies are found in all vegetable and animal
substances; they are not produced by the modes of generation, they
have no uniformity of species, and therefore can neither be animals
nor vegetables. They are to be met with in the flesh of animals, and
in the substance of vegetables, but are most numerous in their seeds;
is it not therefore natural to regard them as living organic particles
which compose the animal or vegetable; as particles which having motion
and a kind of life, ought, by their union, to produce moving and living
beings, and so form animals and vegetables?
But in order to leave this matter as little in doubt as possible,
let us examine other substances. Can it be said, the active machines
which Mr. Needham perceived in the milt of the calmar were animals?
Can it be thought that eggs, which are active machines of another
kind, are also animals? If we turn our eyes to the representation of
almost all the moving bodies Leeuwenhoek saw in different matters,
shall we not be convinced, even at the first inspection, that those
bodies are not animals, since not one of them has any limbs, but
are all either globular or oval? If we afterwards examine what this
celebrated naturalist says, when he describes the motion of these
pretended animals, we can no longer doubt of his being in an error
when he considered them as such; and we shall be still more and more
confirmed that they are only moving organic particles by the following
examples: Leeuwenhoek gives[W] the figure of the moving bodies which he
observed in the liquor of a male frog. This figure only represents a
slender body, long, and pointed at one of its extremities; and of this
he says, "Uno tempore caput (thus he calls the thickest extremity of
this moving body) crassius mihi apparebat alio; plerumque agnoscebam
animalculum haud ulterius quam a capite ad medium corpus, ob caudæ
tenuitatem, & cum idem animalculum paulo vehementius moveretur (quod
tamen tarde fiebat) quasi volumine quodam circa caput ferebatur. Corpus
fere carebat motu; cauda tamen in tres quatuorve flexus volvebatur."
This then is the change of form which I mentioned to have seen, the
mucilage from which the moving bodies use all their efforts to be
disengaged, the slowness of their motion before they are disengaged;
and the animal, according to Leeuwenhoek, one part of which is
in motion, and the other dead: for he afterwards says, "Movebant
posteriorem solum partem, quæ ultima, morti vicinia esse judicabam."
All this does not agree with an animal, but with what I have spoken
of; excepting that I never saw the tail move but by the agitation
of the body. He afterwards says, speaking of the seminal liquor of a
cod, "Non est putandum omnia animalcula in semine aselli contenta uno
eodemque tempore vivere, sed illa potius tantum vivere quæ exitui seu
partui viciniora sunt, quæ & copiosiori humido innatant præ reliquis
vita carentibus, adhuc in crassa materia, quam humor eorum efficit,
jacentibus."
[W] Vol. I. p. 51.
If these are animals, why have they not all life? why are they in the
most fluid part of the liquor alive, while those in the thickest are
not so? Leeawenhoek did not perceive that the thick matter, the origin
of which he attributes to the humour of the animalculæ, is nothing but
a mucilaginous matter which produces them. By diluting this mucilage
with water, he would have given life to the whole of them. Even this
mucilage is oftentimes only a mass of those bodies which are set
in motion on being separated; and consequently this thick matter,
instead of being a humour, produced by the animalcules, is only the
substance of the animals themselves, or rather, as we have already
observed, the matter from which they originate. Speaking of the seed
of a cock, Leeuwenhoek says, in his letter to Grew, "Contemplando
materiam (seminalem) animadverti ibidem tantam abundantiam viventium
animalium, ut ea stuperem; forma seu externa figura sua nostrates
anguillas fluviatiles referebant, vehementissima agitatione movebantur;
quibus tamen substrati videbantur multi & admodum exiles globuli, item
multæ plan-ovales figuræ, quibus etiam vita posset attribui, & quidem
propter earundem commotiones; sed existimabam omnes hasce commotiones
& agitationes pro venire ab animalcules, sicque etiam res se habebat;
attamen ego non opinione solum, sed etiam ad veritatem mihi persuadeo
has particulas planam & ovalem figuram habentes, esse quædam animalcula
inter se ordine suo disposita & mixta vitaque adhuc carentia." Here
we see in the same seminal liquor animalcules of different forms;
and I am convinced, by my own experiments, that if Leeuwenhoek had
closely observed these oval substances, he would have discovered that
they moved by their own powers, and that consequently they were as
much alive as the rest. This change perfectly coincides with what
I have said, that they are organic particles which take different
forms, and not constant species of animals; for in the present case,
if the bodies, which have the figure of an eel, are true spermatic
animalcules, each, destined to become a cock, which supposes a very
perfect organization, and a very constant form, what will those be
which have an oval figure, and what end do they answer? He says indeed
afterwards, that these ovals maybe conceived to be the same animals,
by supposing their bodies to be twisted in a spiral form; but then
how shall we conceive that an animal, whose body is constrained, can
move without being extended? I maintain, therefore, that these oval
substances are no other than the organic particles separated from their
threads, and that the eels were the separated parts which dragged those
threads after them, as I have many times perceived in other seminal
liquors.
Leeuwenhoek, who imagined all these moving bodies were animals, and
established a system thereon; who also pretended, that spermatic
animals must become men and animals, now suspected they were only
natural machines, or organic particles in motion; for he does not doubt
these spermatic animals contained the great animal in miniature, he
says, "Progeneratio animalis ex animalculo in seminibus masculinis
omni exceptione major est; nam etiamsi in animalculo ex semine masculo
unde ortum est, figuram animalis conspicere nequeamus, attamen satis
superque certi esse possumus figuram animalis ex qua animal ortum est,
in animalculo quod in semine masculo reperitur, conclusam jacere sive
esse; & quanquam mihi sæpius conspectis animalculis in semine masculo
animalis, imaginatus fuerim me posse dicere, en ibi caput, en ibi
humeros, en ibi femora; attamen eum ne minima quidem certitudine de iis
judicium ferre potuerim, hujusque certi quid statuere supersedeo, donec
tale animal, cujus semina mascula tam magna erunt, ut in iis figuram
creaturæ ex qua provenit, agnoscere queam, invenire secunda nobis
concedat fortuna." This fortunate chance, which Leeuwenhoek desires,
presented itself to Mr. Needham. Every part of the spermatic animals of
the calmar are easy to be seen without a microscope; but they are not
young calmars, as Leeuwenhoek thinks, nor even animated, although they
are in motion, but only machines which must be regarded as the first
produce of the union of organic particles.
Although Leeuwenhoek had not such an opportunity of undeceiving
himself, he nevertheless had another phenomena which ought to have had
that effect; for example, he had remarked that the spermatic animals of
a dog often change their figures, especially when the liquor was on the
point of evaporating; that these pretended animals had a hole in the
head when they were dead, and that this hole did not appear when they
were alive; he had seen that the part which he looked upon as the head
was full and plump when it was alive, and flaccid and flat when dead.
All this ought to have led him to doubt whether these moving bodies
were real animals; and consider it as agreeing better with a machine,
which empties itself like that of the calmar, than with a moving animal.
I have said that these moving bodies, these organic particles, do
not move like animals, nor have an interval of rest. Leeuwenhoek has
observed the same: "Quotiescunque, says he, animalcula in semine
masculo animalium fucrim contemplatus, attamen illa se unquam ad
quietem contulisse, me nunquam vidisse, mihi dicendum est, si modo
sat fluidæ superesset materiæ in qua sese commode movere poterant;
et eadem in continuo manent motu, & tempore quo ipsis moriendum
appropinquante, motus magis magisque deficit, usquedum nullus prorsus
motus in illis agnoscendus sit." It appears difficult to conceive that
animals can exist, from the moment of their birth till that of their
death, in a continual rapid motion without the least interval of rest;
and I cannot possibly imagine how these animals in the semen of a dog,
which Leeuwenhoek saw the seventh day in as rapid motion as they were
when they were first taken from the body of the animal, preserved
a motion during that time so exceedingly swift, that no animal has
sufficient power to move in for an hour; especially if we consider the
resistance which proceeds from the density and the tenacity of the
liquor. This kind of continued motion, on the contrary, agrees with
the organic particles, which, like artificial machines, produce their
effects in a continual operation, and which stop when that effect is
over.
Among the great number of Leeuwenhoek's experiments, he, without
doubt, often perceived spermatic animals without tails; and he
endeavours to explain this phenomena by a supposition; for example,
he says, speaking of the semen of a cod, "Ubi vero ad lactum accederem
observationem, in iis partibus quas animalcula esse censebam neque
vitam neque caudam dignoscere potui; cujus rei rationem esse
existimabam, quod quamdiu animalcula natando loca sua perfecte mutare
non possunt tam diu etiam cauda concinne circa corpus maneat ordinata,
quodque ideo singula animalcula rotundum repræsentent corpusculorum."
It would have been better to have said, as it in fact is, that the
spermatic animals of these fish have tails at certain times and none
at others, than to suppose their tails twisted so exactly round their
bodies as to give them the shape of a globule. But this must not lead
us to think that Leeuwenhoek only attended to the moving bodies which
he saw with tails, but rather that he did not describe the others,
because, although they were in motion, he did not regard them as
animals; and this is the cause that all the spermatic animals he has
depicted resemble each other, and drawn with tails, since he only
took them for real animals in that state; and that when he saw them
under other forms, he thought them imperfect, or rather that they
were dead. On the whole it appears, by my experiments, that far from
displaying their tails the more as they are in a more perfect condition
of swimming, as Leeuwenhoek says, they, on the contrary, lose their
tails in a gradual manner, till at last these tails, which are no more
than foreign bodies of the animalcules, and which they drag after them,
entirely disappear.
In another part Leeuwenhoek, speaking of the spermatic animals of man,
says, "Aliquando etiam animadverti inter animalcula particulas quasdam
minores & subrotundas; cum vero se ea aliquoties eo modo oculis meis
exhibuerint, ut mihi imaginarer eas exiguis instructas esse caudis,
cogitare coepi annon hæ forte particulæ forent animalcula recens
nata; certum enim mihi est ea etiam animalcula per generationem
provenire, vel ex mole minuscula ad adultam procedere quantitatem: &
quis sit annoa ea animalcula, ubi moriuntur, aliorum animalculorum
nutritioni atque augmini inserviant?" By this passage it appears that
Leeuwenhoek had seen animals without tails in the seminal liquor of
a man, and that he is obliged to suppose them to be just born, and
not adult; but I have observed quite the contrary; for the moving
bodies are never larger than when they separate from the filaments,
and begin to move. When they are entirely disengaged from the mucilage
they become smaller, and continue decreasing as long as they remain
in motion. With respect to the generation of these animals, which
Leeuwenhoek speaks of as certain, I am persuaded no sign of generation
has ever been discovered; all he says is advanced on mere suppositions,
which it is easy to prove by his own observations; for example, he
says that the milt of certain fish, as the cod, fills by degrees with
seminal liquor, which after the fish has emitted, the milt dries up,
leaving only a membrane destitute of any liquor. "Eo tempore, says
he, quo ascellus major lactes suos emisit, rugæ illæ, seu tortiles
lactium partes, usque adeo contrahuntur, ut nihil præter pelliculas
seu membranæ esse videantur." How then does he understand that this
dry membrane, in which there is no longer either seminal liquor or
animalcules, can reproduce animals of the same kind the succeeding
year? if there was a regular generation in these animals, there could
not be this interruption, which in most fishes lasts for a whole year.
To draw himself out of this difficulty, he says, "Necessario statuendum
erit, ut ascellus major semen suum emiserit, in lactibus etiamnum
multum materiæ seminalis gignendis animalculis aptæ remansisse, ex qua
materia plura oportet provenire animalcula seminalia quam anno proxime
elapso emissa fuerant." This supposition, that there remains something
in the seminal liquor in the milts to produce spermatic animals for
the succeeding year, is absolutely contrary to observations, for the
milt is in this interval only a thin and absolutely dry membrane. But
what reply can be made to a still further opposition to this point,
there being fish like the calmar, the seminal liquor of which is not
only renewed every year, but even the reservoir which contains it? Can
it be said, that there remains a seminal matter in the milt for the
production of the animals for the succeeding year, when even the milt
does not remain? it is therefore very certain that these pretended
spermatic animals are not multiplied, like other animals, by the mode
of generation; which alone is sufficient to make us presume, that those
particles which move in the seminal liquors are not real animals. Thus
Leeuwenhoek, who in the passage above quoted says, it is certain that
spermatic animals multiply and propagate by generation, nevertheless
owns, in another part, that the manner in which these animals are
produced is very obscure, and that he leaves to others the task of
clearing up this matter. "Persuadebam mihi," says he, speaking of the
spermatic animals of the dormouse, "hæcce animalcula ovibus prognasci,
quia diversa in orbem jacentia & in semet convoluta videbam; sed unde,
quæso, primam illorum originem derivabimus? in animo nostro concipiemus
horum animalculorum semen jam procreatum esse in ipsa generatione,
hocque semen tam diu in testiculis hominum hærere, usquedum ad annum
ætatis decimum-quartum vel decimum-quintum aut sextum pervenerint,
eademque animalcula tum demum vita donari vel in justam staturam
excrevisse, illoque temporis articulo generandi maturitatem adesse!
sed hæc lampada aliis trado." I do not think it necessary to make any
remarks on what Leeuwenhoek says on this subject: he saw spermatic
animals without tails, and round, in the seed of a dormouse; "in semet
convoluta," says he, because he supposes that they should have tails,
and instead of being certain, as he before had been, that the animals
propagate by generation, he here seems convinced of the contrary. But
when he had observed the generation of pucerons, and was assured[X]
that they engendered without copulation, he caught the idea to explain
the generation of spermatic animals. "Quemadmodum, says he, animalcula
hæc quæ pediculorum antea nomine designavimus (the pucerons) dum adhuc
in utero materno latent, jam prædita sunt materia seminali ex qua
ejusdem generis proditura sunt animalcula, pari ratione cogitare licet
animalculæ in seminibus masculinis ex animalium testiculis non migrate
seu ejici quin post se relinquant minuta animalcula aut saltem materiam
seminalem ex qua iterum alia ejusdem generis animalcula proventura
sunt idque absque coitu; eadem ratione qua supradicta animalcula
generari observavimus." This supposition gives no more satisfaction
than the preceding: for we do not understand by this comparison of
the generation of these animalcules with that of a puceron, why they
are not found in the seminal liquor of a man, before he has attained
the age of fourteen or fifteen years; nor do we know from whence
they proceed, nor how they are renewed every year in fish, &c. and
it appears, that whatever efforts Leeuwenhoek made to establish the
generation of spermatic animals on some probability, it still remained
an entire obscurity, and would, perhaps, perpetually have remained so,
if the preceding experiments had not evinced that they are not animals,
but moving organic particles contained in the nutriment the animal
receives, and which are found in great numbers in the seminal liquor,
which is the most pure, and in the most organic extracts drawn from
this nutriment.
[X] See vol. II. page 499, and vol. III. page 271.
Leeuwenhoek acknowledges that he had not always found animalcules
in the seminal liquor of males; in that of the cock, for example,
which he had often examined, he saw spermatic animals in the form
of eels but once, and some years after he could not discover any
under that form, but observed some with large heads and tails, which
his designer could not perceive. He says also, that one season he
could not find living animals in the seminal liquor of the cod. All
these disappointments proceeded from his desire of finding tails to
these animals; and although he perceived little bodies in motion, he
did not consider them as animals, because they were without tails,
notwithstanding it is under that form they are generally seen, either
in seminal liquors, or infusions of animal or vegetable substances. He
says, in the same place, that he was never able to make his designer
perceive the spermatic animalcules of a cod, which he had so often
seen himself.--"Non solum, says he, ob eximiam eorum exilitatem, sed
etiam quod eorum corpora adeo essent fragilia, ut corpuscula passim
dirumperentur; unde factum fuit ut nonnisi rare, nec sine attentissima
observatione, animadverterem particulas planas atque ovorum in morem
longas, in quibus ex parte caudas dignoscere licebat; particulas
has oviformes existimavi animalcula esse dirupta, quod particulæ hæ
diruptæ quadruplo fere viderentur majores corporibus animalculorum
vivorum." When an animal of any kind ceases to live, it does not then
suddenly alter its form, and from being long, like a thread, becomes
round like a ball; neither does it become four times larger after
its death than it was before. Nothing that Leeuwenhoek says here
agrees with the nature of animals; but, on the contrary, the whole
corresponds with a kind of machine, which, like those of a calmar,
empty themselves after having performed their functions. But let us
pursue this observation; he says, he has seen the spermatic animals
of the cod in different forms, "multa apparebant animalcula sphæram
pellucidam representantia;" he has also seen them of different sizes,
"hæc animalcula minori videbantur mole, quam ubi eadem antehac in tubo
vitreo rotundo examinaveram."
There needs nothing more to shew that there are no constant and
uniform species of these animalcules; and that consequently they are
not animals, but only organic particles in motion, which, by their
different combinations, take different forms and sizes. These organic
moving particles are found in great quantities in the extract and
residue of our nutriment. The matter which adheres to the teeth, and
which in healthy people has the same smell as the seminal liquor, is
only a residue of the food, and a great number of these pretended
animals are also found there, some of which have tails, and resemble
those in the seminal liquor. Mr. Baker had four different kinds of
them engraved, and which were all of a cylindrical or oval make, or
globules with and without tails. I am persuaded, after having strictly
examined them, that not any of them are real animals, but are like
those in the seed, only living organical parts of the nutriment which
present themselves under different forms, Leeuwenhoek, who did not
know how to account for these pretended animals in the matter which
adhered to the teeth, supposed them to proceed from certain food they
were previously in, as cheese, &c. but we find them among the teeth of
those who do not eat cheese, as well as in those that do; besides, they
have not the least resemblance to mites, nor the other animalcules seen
in rotten cheese. In another place he says, these animals of the teeth
may proceed from the cistern water that is drank, because he observed
animals like them in dew and rain water, especially in that which
stagnates upon lead and tiles; but with which we can prove there is not
the least resemblance.
Most seminal liquors dilute of themselves, and liquefy when exposed to
the air or a certain degree of cold; but they thicken when a moderate
degree of heat is communicated to them. I have exposed some of these
liquors to a very intense cold, as water on the point of freezing, but
it did no injury to these supposed animals; they continued to move with
the same swiftness, and as long as those which had not been so exposed,
but those which had suffered but a little warmth soon ceased to move,
because the liquor thickened. If the moving bodies were animals,
they were of a complexion and temperament quite different from all
others, to whom a gentle and moderate heat strengthens their powers and
motions, which the cold stops and destroys.
Notwithstanding it may be thought I have dwelt too long upon this
subject, I cannot conclude it without making one remark, from which
some useful conclusions may be drawn. These pretended spermatic
animals, which are only living organic particles of the nutriment, not
only exist in the seminal liquors of the two sexes, and in the residue
of the nutriment which adheres to the teeth, but also in the chyle
and excrements. Leeuwenhoek having met with them in the excrements
of frogs, and other animals, which he dissected, was at first very
much surprised, and notable to conceive from whence these animals
proceeded, so entirely like those he had observed in the seminal
liquors, accuses himself of having, in dissecting the animal, opened
the seminal vessels, and that the seed had by that means been mixed
with the excrements. But having afterwards found them in the excrements
of other animals, and even in his own, he no longer knew to what to
attribute them. Leeuwenhoek, it is worthy remark, never met with
them in his own excrements, but when they were liquid. Every time he
was disordered and the stomach did not perform its functions, and was
relaxed, he discovered these animalcules; but when the concoction of
the food was well performed, and the excrement was hard, there was not
a single one, although it was diluted with water. This seems perfectly
to agree with all we have before advanced: for when the stomach and
intestines perform their functions, the excrements are only the grosser
parts of the nutriment; and all that is really nutritive and organic
passes into the vessels which serve to nourish the animal; whereas if
the stomach and intestines are not in a condition to comminute the
food, then it passes with the inanimate parts, and we find the living
organic molecules in the excrements; from whence it may be concluded,
that those which are often lax must have less seminal liquor, and be
less proper for generation, than those of a different habit of body.
In all I have said, I constantly supposed the female furnished a
seminal liquor, which was as necessary to generation as that of the
male. I have endeavoured to establish in Chap. I. that every organized
body must contain living organic particles, and I have endeavoured to
prove Chap. II. and III. that nutrition and reproduction operates by
the same cause; that nutrition is made by the intimate penetration
of these organic particles through each part of the body, and that
reproduction operates by the superfluity of these same organic
particles collected together from all parts of the body and deposited
in proper reservoirs. I have explained in Chap. IV. how this theory
must be understood in the generation of man and animals which have
sexes. Females then being organized bodies like males, they must also
have some reservoirs for the superfluity of organic particles returned
from every part of their bodies. This superfluity cannot come there
through any other form than that of a liquor, since it is an extract of
all parts of the body; and this liquor is that to which I have given
the name of the female semen.
This liquor is not, as Aristotle pretends, an infecund matter of
itself, which enters neither as matter nor form into the business
of generation, but as essentially prolific as that of the male,
containing characteristic parts of the feminine sex, which the female
alone can produce, the same as the male contains particles necessary
to form the masculine organs; and each of them contains every other
organic particle that can be looked on as common to both sexes; which
causes that, by their mixture, the daughter may resemble her father,
and the son his mother. This semen Hippocrates says, is composed of
two liquors; the one strong, for the production of males; and the
other weak, for the production of females. But this supposition is
too extended; I do not see how it is to be conceived that a liquor,
which is the extract of every part of the female body, should contain
particles for the formation of the male organs.
This liquor must enter by some way into the matrix of animals which
bear and nourish their foetus within the body, and in others, as
oviparous animals, it must be absorbed by the eggs, which may be looked
upon as portable matrixes. Each of these matrixes contains a small
drop of this prolific liquor of the female, in the part that is called
the _cicatrice_. When there has been no communication with the male,
this prolific drop collects under the form of a small mole, or mass,
as Malpighius observes; but when impregnated by that of the male; it
produces a foetus which receives its nutriment from the juices of the
egg.
Eggs, instead of being parts generally found in every female, are
therefore only instruments made use of by Nature to serve as the matrix
in females which are deprived of that organ. Instead also of being
active and essential to the first fecundation, they only serve as
passive and accidental parts for the nutrition of the foetus already
formed by the mixture of the liquor of the two sexes in a particular
part of this matrix. Instead also of being existing bodies, inclosed,
_ad infinitum_, one within the other, eggs, on the contrary, are bodies
formed from the superfluity of a more gross and less organic part of
the food, than that which produces the seminal and prolific liquor; and
are in oviparous females something equivalent, not only to the matrix,
but even to the menstrua in the viviparous.
We should be perfectly convinced, that eggs are only destined by Nature
to serve as a matrix in animals who have not that viscera, by those
females producing eggs independant of the male. In the same manner as
the matrix exists in viviparous animals, as a part appertaining to the
female sex, hens, which have no matrix, have eggs in their room, which
are successively produced of themselves, and necessarily exist in the
female independently of any communication with the male. To pretend
that the foetus is pre-existing in the eggs, and that these eggs are
contained, _ad infinitum_, within each other, is nearly the same as
to pretend that the foetus, is pre-existing in the matrix, and that
the matrix of the first female inclosed all that ever were or will be
produced.
Anatomists have taken the word _egg_ in several acceptations and
meanings. When Harvey took for his motto, _Omnia ex ovo_, he understood
by the word egg, as applied to viviparous animals, the membrane which
includes the foetus and all its appendages: he thought, he perceived
this egg, or membrane, form immediately after the copulation of the
male and the female. But this egg does not proceed from the ovium
of the female; and he has even maintained, that he did not remark
the least alteration in this testicle, &c. We perceive there is here
nothing like what is commonly understood by the word egg unless the
figure of the bag may be supposed to have some resemblance thereto.
Harvey, who dissected so many viviparous females, did not, he says,
ever perceive any alteration in the ovaria; he looked on them even
as small glands, perfectly useless to general ion,[Y] although they
undergo very remarkable changes and alterations in them, since we
may perceive in cows the glandular bodies grow from the size of a
millet seed to that of a cherry. This great anatomist was led into
this error by the smallness of the glandular bodies in the species
of deer, to which he principally paid his attention. C. Peyerus, who
also made many experiments on them, says, "Exigui quidem sunt damarum
testiculi, sed post coitum foecundum, in alterutro eorum, papilla,
sive tuberculum fibrosum, semper succrescit; scrofis autem prægnantibus
tanta accidit testiculorum mutatio, ut mediocrem quoque attentionem
fugere nequeat."[Z] This author imagines, with some reason, that the
minuteness of the testicles of does, is the cause of Harvey's not
having remarked the alterations; but he is wrong in advancing that the
alterations he had remarked, and which had escaped Harvey's notice, did
not happen till after impregnation.
[Y] See Harvey Exercit. 64 and 65.
[Z] Vide Conradi Peyeri Merycologia.
It appears that Harvey was deceived in many other essential points;
he asserts, that the seed of the male does not enter into the matrix
of the female, and even that it cannot; yet Verheyen found a great
quantity of the male seed in the matrix of a cow, which he dissected
six hours after copulation.[AA] The celebrated Ruysch asserts, that
having dissected a woman who had been caught in the act of adultery,
and was assassinated, he found, not only in the cavity of the matrix,
but also in the trunks, a quantity of the seminal liquor of the
male,[AB] Valisnieri affirms, that Fallopius and other anatomists
had also discovered male seed in the matrix of many women. After the
positive testimony of these great anatomists, there can remain no doubt
but Harvey was deceived in this important point; especially when to
these are added that of Leeuwenhoek, who found the male seed in the
matrix of a great number of females of different species.
[AA] See Verheyen Sup. Anat. Tra. v. cap. iii.
[AB] See Ruysch, Thes. Anat. p. 90, tab. VI, fig. I.
Harvey makes another error in speaking of an abortion in the second
month, where the mass was as large as a pigeon's egg, but without any
foetus regularly formed; whereas, it is maintained by Ruysch, and
many other anatomists, that the foetus is perceptible, even to the
naked eye, in the first month. The History of the Academy mentions
a foetus, that was completely formed in twenty-one days after
impregnation. If to these authorities we add that of Malpighius,
who perceived the chicken in the cicatrice, immediately after the
egg was laid by the hen, we cannot doubt, but that the foetus is
formed immediately after copulation; consequently, we must not pay any
credit to what Harvey says on the parts increasing one after the other
by juxta-position, since they are all existent from the first, and
gradually expand until the whole is complete.
De Graaf took the acceptation of the word egg in a quite different
light to Harvey: he insists that the testicles of women were true
ovaries, and contain eggs like those of oviparous, animals, only that
they are much smaller, do not quit the body, and are never detached
till after impregnation, when they descend from the ovary into the
horns of the matrix. The experiments of De Graaf have contributed most
to establish the existence of these pretended eggs, which yet is not at
all founded; for this famous anatomist is deceived, first, by mistaking
the vesicles of the ovarium for eggs, whereas they are inseparable from
it, form parts of its substance, and are filled with a kind of lymph.
Secondly, he is also deceived when he considers the glandular bodies
to be the covering of those eggs, or vesicles; for it is certain,
by Malpighius's, Valisnieri's, and my own observations, that the
glandular bodies neither surround nor contain one of those vesicles.
Thirdly, he is deceived still more when he supposes the glandular body
is never formed till after fecundation; as they are invariably found in
every female who has attained the age of puberty. Fourthly, he is no
less deceived when he believes that the globules which he saw in the
matrix, and which contained the foetuses, ware the same vesicles,
or eggs, which had fallen from the ovariam, and which, he remarks,
were become ten times smaller than they were in the ovary. This remark
alone, one would imagine, Should have made him perceive his error.
Fifthly, he is wrong in saying that the glandular bodies are only the
coverings of the fecundated eggs, and that the number of coverings,
or empty follicles, always answer to the number of foetuses. This
assertion is entirely contrary to truth: for on the testicles of all
females we find a greater number of glandular bodies, or cicatrices,
than there are productions of foetuses, and they are also found in
those which have never brought forth. To this we may add, that neither
he, Verheyen, nor any other person, have ever seen these eggs, much
less these pretended coverings, on which they have, notwithstanding,
established their system.
Malpighius, who perceived the growth of the glandular bodies in the
female testicles, was deceived when he thought he had seen the egg in
their cavities, since they contain only liquor; nor indeed has anything
like an egg ever been discovered.
Valisnieri, who was not deceived in facts, has yet drawn false
conclusions in asserting that, although neither himself, nor any
anatomist in whom he could confide, ever found the egg in the cavity of
the glandular body, yet it must there exist.
Let us, therefore, examine what may be fairly called the real
discoveries of these naturalists. Graaf was the first who perceived
there were alterations in the female testicles; and he had reason
to affirm, they were parts essential and necessary to generation.
Malpighius demonstrated that these alterations were occasioned by the
glandular bodies which grew to perfect maturity, afterwards they become
flaccid, obliterated, and left only a slight cicatrice remaining.
Valisnieri has placed this discovery in a very clear light; he has
shewn that these glandular bodies are found in the testicles of every
female; that they are augmented considerably in the season of love,
that they increase at the expence of the lymphatic vesicles of the
testicles, and that at the time of their maturity they were hollow and
filled with liquor. This, then, is all that can be reduced to truth on
the subject of the pretended ovaries and eggs of viviparous animals.
What must we conclude therefrom? Two things appear very evident: the
one, that there does not exist any eggs in the female testicles; the
other, that there exists a liquor in the vesicles of the testicle, and
in the cavity of the glandular bodies. We have demonstrated by the
preceding experiments, that this last liquor is the true seed of the
female, since it contains, like that of the male, spermatic animals, or
rather organic moving particles.
We must, therefore, now be assured, that females have, as well as
males, a seminal liquor. After all that has been advanced, we cannot
doubt but the seminal liquor is the superfluity of the organic
nutriment, which is sent back from all parts of the body into the
testicles and seminal vesicles of the males, and into the testicles
and glandular bodies of females. This liquor, which issues by the
nipple of the glandular bodies, continually sprinkles the horns of the
matrix, and may easily procure admission either by the suction of the
membrane of these horns, or by the little opening which is at the upper
extremity, and thus enter into the matrix; but in the supposition of
these pretended eggs, which were ten or twenty times larger than the
opening of the horns of the matrix, we cannot comprehend how they could
enter therein.
The liquor emitted by females, when they are excited, and which,
according to de Graaf, issues from the neck of the matrix, and the
orifice of the urethra, may be a superabundant portion of the seminal
liquor which continually distills from the glandular bodies on the
trunks of the matrix. But, possibly, this liquor may be a secretion
of another kind, and perfectly useless in generation. To decide this
question observations with a microscope are requisite; but _all_
experiments are not permitted even to philosophers. I can only say,
that I am inclined to believe that the same spermatic animals would
be met with in this liquor as in that of the glandular bodies. I can
quote an Italian doctor on this subject, who made this observation with
attention, and which is thus related by Valisnieri: "Aggiugne il lodato
fig. Bono d'avergli anco veduti (animali spermatici) in questa linfa o
siero, diro cosi voluttuoso, che nel tempore dell'amorosa zuffa scappa
dalle femine libidinose, senza che si potesse sospettare che fossero
di que' del maschio, &c." If this circumstance is true, as I do not
doubt, it is certain, that this liquor is the same as that found in
the glandular bodies, and that, consequently, it is the true seminal
liquor: and although anatomists have not discovered the communication
between the vacuities of de Graaf and the testicles, that does not
prevent it being once in the matrix, from issuing out by the vacuities
about the exterior orifice of the urethra.
From hence we must conclude that the most abandoned women will be the
least fruitful, because they emit that liquor which ought to remain in
the matrix for the formation of the foetus. Thus we see why common
prostitutes seldom have children, and why women in hot countries, where
they have stronger desires than in the cold, are much less fertile; but
we shall have occasion to speak of this hereafter.
It is natural to think that the seminal liquor of the male or female
would not be fertile but when it contains moving bodies; nevertheless
that is still a question, and I should be led to think, as there are
different states of this liquor, that in which these organic particles
are seen in motion is not absolutely necessary for the purpose of
generation. The Italian physician, above quoted, never perceived
spermatic animals in his semen till he had attained a middle age,
although he was father of several children before, and continued to
have them afterwards.
These spermatic bodies, which move, may be looked upon as the first
assemblages of the organic molecules which proceed from every part of
the body; when a quantity of them collect they may be perceived with
the microscope; but if they collect only in small quantity the body
which they form will be too minute to be perceived, and in this case we
shall not be able to distinguish any in the seminal liquor. A very long
continuance of observations would be necessary to determine what can be
the cause of all the differences remarked in the states of this liquor.
I can assert, from having often tried it, that by infusing the seminal
liquors in water closely corked, at the end of three or four days
an infinite multitude of moving bodies will be found, although the
seminal liquors had no motion on being first taken from the body of
the animal. Flesh, blood, chyle, urine, nay all animal or vegetable
substances, contain organic particles, which move at the end of some
days in an infusion of water; they appear to act and move nearly in the
same manner, and though produced from different bodies are perfectly
similar, without any of them having a power peculiar to themselves.
If these bodies must absolutely be termed animals, it must be allowed
they are so imperfect that they ought to be looked upon as the outlines
of them, or rather as bodies simply composed of particles the most
essential to the existence of an animal; for natural machines, such as
those found in the roe of a calmar, although they put themselves in
action at certain times, are certainly not animals, although they are
organized, acting, and, as I may say, living beings.
If it is once allowed, that the productions of Nature follow in an
uniform order, and advance by imperceptible degrees and links, we shall
have no difficulty in conceiving there are organic bodies existing,
which belong neither to animals, vegetables, nor minerals.
It is certain, however, that all animals and vegetables contain an
infinity of organic living molecules. These molecules successively
take different forms, and different degrees of motion and activity,
according to different circumstances They are in a much greater number
in the seminal liquor of both sexes, and in the germs of plants, than
in other parts of the animal or vegetable. There exists, then, a
living substance in animals and vegetables, common to both, and which
substance is necessary to their nutrition. An animal procures nutriment
from an animal or vegetable substance, and the vegetable can likewise
be nourished from an animal or vegetable in a decomposed state. This
nutritive substance, common to both, is always living, always active,
and produces an animal or vegetable, as it finds an internal mould or
an analogous matrix, as we have explained in the first chapters; but
when this active substance collects in great abundance, in those parts
where it can unite, it forms in the animal body other living creatures,
such as the tape-worm, ascarides, and worms, which are sometimes found
in the veins, in the sinus of the brain, in the liver, &c. These kinds
of animals do not owe their existence to the animals of the same
species, and we may, therefore, suppose, they are produced by this
organic matter when it is extravasated, or is too abundant for the
lacteal vessels to absorb. We shall hereafter have occasion to examine
more largely the nature of those worms, and many other animals which
are formed in a similar manner.
When this organic matter, which may be looked on as an universal seed,
is collected in any great quantity, as in the seminal liquors, and in
the mucilaginous parts of the infusion of plants, its first effect is
to vegetate, or rather to produce vegetating beings. These zoophytes
swell, extend, ramify, and produce globules, ovals, and other small
bodies, of different figures, which have all a kind of animal life, a
progressive motion, which is often very swift, and sometimes very slow.
These globules themselves decompose, change their figures, and become
smaller; and in proportion as they diminish in size the rapidity of
their motion augments.
I have sometimes thought that the venom of the viper, and other active
poisons, even that of the bite of a mad dog, might possibly be this
active matter too rarefied; but I have not as yet had time to make the
experiments which I had projected on this matter, as well as on drugs
used in medicine; all that I can at present ascertain is, that all
infusions of the most active drugs swarm with moving bodies, which form
therein in much less time than in other substances.
Almost all microscopic animals are of the same nature as the organized
bodies which move in the seminal liquor, in the infusions of vegetables
and the flesh of animals; the eel-like bodies in flour, vinegar, and
water, in which lead has been soaked, are beings of the same nature as
the first, and have a like origin.
CHAPTER IX.
VARIETIES IN THE GENERATION OF ANIMALS.
The matter which serves for the nutrition and reproduction of animals
and vegetables is therefore the same; it is a productive and universal
substance, composed of organic molecules, and whose union produces
organized bodies. Nature always works on the same fund, which is
inexhaustible, but the means she employs to stamp its value are
different, and these differences, or general agreements, deserve
attention, because it is from thence we must derive our reasons to
account for exceptions and particular varieties.
In general large animals are less productive than small. The whale,
elephant, rhinoceros, camel, horse, the human species, &c. only
produce one, and very seldom two, at a birth; whereas small animals,
as rats, herrings, insects, &c. produce a great number at a time. Does
not this difference proceed from there being more food required to
support a large body than to nourish a small one, and from hence the
former has less superfluous organic particles, which would convert
into semen, than the latter? It is certain that small animals eat more
in proportion than large ones; but it is likewise probable that the
prodigious multiplication of the small animals, as bees, flies, and
other insects, may be attributed to their being endowed with very fine
and slender limbs and organs, by which they are in a condition to chuse
what is most substantial and organic in the vegetable or animal matters
from whence they derive their nutriment. A bee, who lives on the
purest parts of flowers, certainly receives more organic particles in
proportion than a horse who feeds on the grosser parts of vegetables,
hay, &c. The horse produces but one at one time, whereas the bee will
bring forth three thousand.
Oviparous animals are in general smaller than the viviparous, and
produce also more at a birth. The duration of the foetus in the
matrix of viviparous animals likewise opposes their increase, nor can
there be any new generation take place during gestation, or while they
are suckling their young; whereas oviparous animals produce at the same
time both matrix and foetuses, which they cast out of the body, and
are therefore almost always in a state of reproduction; and it is well
known that by preventing a hen from setting, and largely feeding, the
number of her eggs will be considerably increased. If hens cease to
lay when they sit, it is because they have ceased to feed; and it is
the fear lest their eggs should not produce which causes them not to
quit their nests but once a day, and that for a very short time, during
which they take a little nutriment, but not one-tenth part of what they
take at other times.
Animals which produce but a small number at a time, acquire the chief
part of their growth before they are fit for engendering, whereas those
which multiply numerously generate before they have received half their
growth. The human species, the horse, the ass, the goat, and the ram,
are not able to engender until they have obtained nearly the whole of
their growth. It is the same with pigeons and other birds, who lay but
a few eggs; but those which produce in great numbers, as poultry, fish,
&c. engender much sooner. A cock is capable of engendering at the age
of three months, when he has not attained a third part of his growth;
a fish, which at the end of twenty years will weigh thirty pounds,
engenders in the first or second year, when perhaps it does not weigh
half a pound. But exact observations on the growth and duration of
the life of fish are still wanting: their age may be nearly known by
examining the annual layers of their scales; but we are not certain how
far that may extend. I have seen carp in the Comte de Maurepas' canals,
at his castle at Pont Chartrain, which were said to be 150 years old,
and they appeared as brisk and lively as the common carp. I will not
say, with Leeuwenhoek, that fish are immortal, or at least can never
die with age; all must perish in time, that is; all which have a
beginning, a birth, must arrive to an end, or death; but fish, living
in an uniform element, and being sheltered from the vicissitudes and
all the injuries of the air, must live a longer time in the same state
than other animals, especially if these vicissitudes of the air be, as
a great philosopher asserts, the principal causes of the destruction of
living beings. But what must contribute to the long duration of their
life is, that their bones are softer than those of other animals, and
do not harden with age. The bones of fish lengthen, and grow thick
without taking any more solidity; whereas the bones of other animals
continually increase in hardness and density, until at length, being
absolutely full, the motion of their fluid ceases, and death ensues. In
their bones the repletion or obstruction, which is the cause of natural
death, is formed by such slow and insensible degrees, that fish must
require much time to arrive at what we call old age.
All quadrupeds covered with hair are viviparous; all those covered
with scales oviparous. May we not then believe than in oviparous
quadrupeds, a much less waste is made by transpiration, than the
cloathing of scales retains; whereas in animals covered with hair
this transpiration is more free and abundant? and is it not partly
by this superabundance of nutriment, which cannot be carried off by
transpiration, that those animals multiply so abundantly, and are
enabled to go so long without food? All birds and all insects that fly
are oviparous, excepting some kinds of flies which bring forth their
young alive. These flies have no wings at their birth, but they shoot
out and grow by degrees, and which they cannot use before they are of
full growth. Scaly fish are likewise oviparous; as are all reptiles
which have no legs, such as snakes and different kinds of serpents;
they change their skins, which are composed of small scales. The viper
is only a slight exception to the general rule, for it is not truly
viviparous, as it produces eggs, from which the young are hatched; it
is certain this is performed in the body of the mother, who instead
of casting those eggs, like other oviparous animals, she retains and
hatches them in her own body. The salamander, in which eggs and young
ones are found at the same time, as observed by M. de Maupertuis, is
an exception of the same kind in oviparous quadrupeds.
Most animals are perpetuated by copulation; yet many birds seem only
strongly to compress the females; indeed the ostrich, Crane, and
some few others, are so well supplied as to leave intromission no
ways equivocal. Male fish approach the female in the spawning time;
they seem even to rub their bellies against each other, for the male
often turns upon its back to meet the belly of the female; but the
necessary part for copulation does not exist in them; and the male
fish approaches the female only to emit the liquor in their milts on
the eggs, which the female then deposits; and it seems rather to be
attracted by the eggs than the female; for when she ceases throwing out
the eggs, he instantly forsakes her, and with eagerness pursues the
eggs, which the stream carries away, or that the wind disperses. Male
fish may be seen to pass and repass every spot where eggs are deposited
several times. It is certainly not for the love he bears the female
that all these motions are made, because it is not to be presumed he
always knows her; often being seen to emit his liquor on all eggs that
he comes near, and that often before he has met with the female to
which they belonged.
There are therefore animals, distinguished by sexes, which have proper
parts for copulation, and some which are deficient in them; others, as
snails, have both, and the two sexes in the same individual; others,
as vine-fretters, have no sex, and engender in themselves separately;
although they couple together when they please, we cannot determine
whether that is a conjunction of sexes; if it is so, we must suppose
that Nature has included in this small individual more faculties for
generation than in any other kind of animal, and that it not only has
the power of reproducing distinctly, but also the means of multiplying
by the communication of another individual.
But whatever difference takes place in generation, Nature, by a new
production, prepares the body for it, and which, whether manifested
outwardly, or concealed internally, always precedes generation. The
ovaries of oviparous animals, and the testicles of female viviparous
animals, before the season of impregnation, experience a considerable
change. Oviparous animals produce eggs, which at first are attached to
the ovaries, by degrees they increase in size, until they fall into
the canal of the matrix, where they acquire their white membranes, and
shell. This production has marks of the fecundity of the female, and
without which generation cannot be performed: so in viviparous females
there are always one or more glandular bodies on the testicles, which
by degrees grow under the membrane that surrounds them; these glandular
bodies enlarge and pierce, or rather impel and lift up the membrane of
the testicle; when their maturity is complete, a small slit or several
small holes appear at their extremities, by which the seminal liquor
escapes, and falls into the matrix: these glandular bodies are new
productions that precede generation, and without which there would not
be any.
In males there is also a similar change which always precedes their
capacity for generating. In oviparous animals a great quantity of
liquor fills a considerable reservoir, and which reservoir itself is
sometimes formed every year; as in the calmar and some other fish. The
testicles of birds swell surprisingly just preceding their amorous
season. In viviparous males the testicles also swell considerably in
those who have seasons, and in general there is a swelling and an
extension of the genital members in all species, which, although it be
external, must be regarded as a new production necessarily preceding
generation.
In the body of every animal, male or female, new productions are formed
which precede generation; and when there is no real production there
is always a swelling, and considerable extension in some of the parts.
There are species in which this new production is not only manifest,
but even the whole body seems to be renewed before generation can be
performed; as is the case with insects whose various metamorphoses
seem to be only for the purpose of generating; for the growth of the
animal is completed before it is transformed. It ceases from taking
nutriment, has no organs for generation, no means of converting the
nutritive particles, of which they abound, into eggs or seminal liquor,
and therefore this superfluity unites and moulds itself at first into
a form something like that of the original. The caterpillar becomes
a butterfly, because, for these reasons, it is unable to produce
small organized beings like itself; the organic particles, always
active, take another form, by uniting, whose figure answers in part,
and even in essential constitution, to that of the caterpillar, but
in which the organs of generation are developed, and may receive
and transmit the organic particles of the nutriment which forms the
eggs, and the individuals of the species. The individuals which
proceed from the butterfly ought not to be butterflies, because the
nutriment, from whence the organic particles proceed, was taken while
in the form of caterpillars; the produce therefore must be similar,
and not butterflies, which is only an occasional production of the
superabundant nutriment; a method adapted by Nature to accomplish the
purposes of generation in these species, as by the glandular bodies and
milts in other animals.
When the superabundant quantity of organic nutriment is not great,
as in man and most large animals, generation is not made till the
growth of the animal is nearly complete, and then it is confined to
the production of a small number of individuals. When these particles
are more abundant, as in many kinds of birds, and in oviparous fishes,
generation is completed before the animal has received its full
growth, and their production of individuals is very numerous. When
the quantity of particles is still greater, as in insects, it first
forms a large organic body, which, though retaining the essential
constitution of its original, differs in many parts, as the butterfly
from the caterpillar, but shortly produces an astonishing number of
young, similar in form to the animal which selected the nutriment. When
the superabundance is greater still, and when at the same time the
animal has the necessary organs for generation, as the vine-fretter, it
immediately produces a generation in every individual, and afterwards
a transformation, like other insects. The vine-fretter becomes a fly,
but cannot produce any thing, because it is only the remainder of the
organized particles which had not been made use of in the production of
the young.
Almost every animal except man has stated times for generation. Spring
is marked out for birds. Carp, and many kinds of fish, spawn in June
and August. Barbel, and other kinds, in spring. Cats have three
seasons, in January, May, and September. Roebucks, in December. Wolves
and Foxes, in January. Horses, in summer. Stags, in September and
October; and almost all insects generate in autumn: these last seem to
be totally exhausted by generation, and die a short time after. Other
animals, though not exhausted, become extremely lean and very weak,
and require a considerable time to repair the loss which is made of
the organic substance. Others are exhausted still less, and are soon
restored to an engendering state; while man is scarcely in the least
affected; his loss is speedily repaired, and therefore may be said to
be at all times in a state for propagation; all which depends solely
on the particular construction of the animal organs. The grand limits
Nature has placed in the mode of existence are equally conspicuous
in the manner of receiving and digesting the food, in the manner of
retaining it in, or excluding it from, the body, and in the means by
which the organic molecules, necessary for reproduction, are extracted.
In a word, we shall find throughout all nature, that all what can be,
is.
The same difference exists in the time of female gestation; some, as
mares, carry their young eleven or twelve months; others, as women,
cows, &c. nine months; others, as foxes, wolves, &c. five months;
bitches, nine weeks; cats, six weeks; rabbits, thirty-one days. Most
birds come out of the egg at the end of twenty-one days; though some,
as canary birds, hatch in thirteen or fourteen days. The variety is
as great here as in every thing else relative to animals. The largest
animals which produce only few, are those which go the longest with
young; this still more confirms what we have already said, that the
quantity of organic food is in proportion less in large than in small
animals; for it is from the superfluity of the mother's food that the
foetus derives what is necessary to the growth and expansion of its
parts, and since this expansion demands much more time in large than
in small animals, it is a proof that the quantity of matter which
contributes is not so abundant in the first as in the last.
There is, therefore, an infinite variety in animals, with respect to
the time and manner of gestation, engendering, and bringing forth; and
this variety is found even in the causes of generation; for although
the general principle of production is this organic matter common to
all that lives or vegetates, the manner in which the union is made,
must have infinite combinations, which must all proceed from the source
of new productions. My experiments clearly demonstrate, that there are
no pre-existing germs, and at the same time prove that the generation
of animals and vegetables is not equivocal; there are, perhaps, as
many beings, either living or vegetating, which are produced by the
fortuitous assemblage of organic molecules, as by a constant and
successive generation. It is to those productions we should apply
the axiom of the ancients, "Corruptio unius, generatio alterius."
The corruption and composition of animals and vegetables produce an
infinite number of organized bodies; some, as those of the calmar, form
only kinds of machines, which, although very simple, are exceedingly
active; others, as the spermatic animalcules, seem by their motion, to
imitate animals; others imitate vegetables by their manner of growing
or extending; there are others, as those of blighted corn, which may
be made to live and die alternately, and as often as we please; there
are still others, even in great quantities, which are at first kinds of
vegetables, afterwards become species of animals, then return again to
vegetables, and so on alternately. There is a great appearance, that
the more we shall observe this race of organized beings, the more we
shall discover varieties, always so much the more singular as they are
the more remote from our sight, and from the varieties of other animals
that have already become known to us.
For example, spurred barley, which is produced by an alteration or
decomposition of the organic substance of the grain, is composed of
an infinity of little organized bodies, like to eels. By infusing
the grain for ten or twelve hours in water, we find them to have a
remarkable twirling, and a slight progressive motion; when almost dry,
they cease to move, but by adding fresh water their motion returns.
The same effects may be produced for months, or even years; insomuch
that we can make these little machines act as often and as long as we
please without destroying them, or their losing any of their power or
activity. Their threads will sometimes open, like the filaments of
semen, and produce moving globules; we may therefore suppose them to be
of the same nature, only more fixed and solid.
Eels, in paste made with flour, have no other origin than the union of
the organic particles of the most essential parts of the grain: the
first which appear are certainly not produced by many others; yet,
although they have not been engendered, they engender others. By
cutting them with the point of a lancet, we may perceive small eels
come from their bodies in great numbers; the body of the animal appears
to be only a sheath or bag which contains a multitude of other little
animals, which perhaps are themselves only sheaths of the same kind, in
which the organic matter assimilates, and takes the form of eels.
There requires a great number of observations to be made to establish
classes and races between such singular beings, which are at present so
little known; there are some which may be regarded as real zoophytes,
which vegetate, and at the same time appear to twirl and move like
animals. There are some that at first appear to be animals, which
afterwards join and form kinds of vegetables. A little attention to the
decomposition of a grain of wheat infused in water will elucidate all I
have asserted. I could add more examples, but I have related these only
to point out the varieties there are in generation. There are certainly
organized beings which we regard as animals, but which are not
engendered by others of the same kind; there are some which are only
a kind of machines, whose action is limited to a certain effect, and
which can act but once in such a certain time, as those in the calmar;
and there are others, as we have just remarked, which we can cause to
act as long and as often as we please. There are vegetating beings
which produce animated bodies, as the filaments of the human seed,
from whence the active globules spring, and which move by their own
powers. In the corruption, fermentation, or rather the decomposition
of animal and vegetable substances, there are organized bodies which
are real animals, and can propagate their like, although they have
not been so produced. The limits of these varieties are perhaps still
greater than we can imagine. We may extend our ideas, and exert every
effort to reduce the effects of Nature to certain points, and class her
productions to certain classes, yet an infinite number of links will
always escape us.
CHAPTER X.
OF THE FORMATION OF THE FOETUS.
It appears to be clearly ascertained by the experiments of Verheyen,
who in one of them found the seed of a bull in the matrix of a cow;
and by those of Ruysch, Fallopius, Leeuwenhoek, and many others, who
perceived the male semen in the uterus of women, and numberless other
animals, that the seminal liquor of the male enters by some means
into the matrix of the female. It is probable, that in the time of
copulation the orifice of the matrix opens to receive the seminal
liquor, but if that is not the case, the active and prolific substance
of this liquor, may penetrate the membranes of the matrix; for the
seminal liquor being, as we have proved, almost all composed of organic
molecules, which are in great motion, and extremely minute, they may
pass across the coat of the closest membranes, and penetrate those of
the matrix with the greatest facility.
What proves that the active part of this liquor may not only pass
through the pores of the matrix, but even penetrate its substance, is
the sudden change that immediately takes place after conception. The
menses are suppressed, the matrix becomes softer, swells, and appears
inflamed. All these alterations can only happen by the action of an
external cause; by the penetration of some part of the seminal liquor
into the substance even of the matrix. This penetration not only
operates on the external surface of the matrix, but on all the other
parts of which this viscera is composed, like that penetration by which
nutrition and expansion is produced.
We shall be easily persuaded that it is so, when we consider that
the matrix, during the time of gestation, not only augments in bulk
but also in quantity of matter, and that it has a kind of life or
vegetation, which is continually increasing till the time of delivery;
for if the matrix was only a pouch, a destined receptacle to receive
the seed and contain the foetus, it would extend and grow thin in
proportion as the foetus increased in size; but in reality the matrix
not only extends in proportion as the foetus grows larger, but
receives at the same time a thickness and solidity. This augmentation
is a real growth, like the expansion of the body in young animals,
which can only be produced by the intimate penetration of the organic
molecules analogous to the substance of the parts: and as this
expansion of the matrix never happens but after impregnation, we cannot
doubt its being produced by the liquor of the male, especially as the
expansion takes place before the foetus has sufficient bulk to dilate
it.
It seems certain, by my experiments, that the female has a seminal
liquor which commences to be formed in the testicles, and is completed
in the glandular bodies: this liquor distills through the small holes,
at the extremities of these bodies; and may, like that of the male,
enter into the matrix in two different manners, either by these holes
at the extremities, or through the membraneous coat of the matrix.
These seminal liquors are both extracts from all parts of the body, and
in the mixture of them there is every thing necessary to form a certain
number of males and females; and the more the animal abounds with this
liquor, and the more that abounds with organic molecules, the greater
is their number of young; as we have already remarked is the case with
the small animals, and diminishes in the large.
But to pursue our subject with greater attention, we shall first
examine the particular formation of the human foetus, and afterwards
return to the other animals. In the human species, as well as in large
animals, the seminal liquors of the male and female do not contain a
great abundance of organic molecules, and therefore seldom produce
more than one at a time: the foetus is a male, if the number of the
organic molecules of the male predominates in the mixture, and a female
if the contrary; and it resembles the father or the mother as they
happen to abound in the mixture of the two liquors.
I conceive, therefore, that the seminal liquor of both are two matters
equally active and necessary for generation; and this I think is
sufficiently proved by my experiments, since I have seen the same
moving bodies in the one as the other. I perceived that the liquor
of the male enters into the matrix, where it meets with that of the
female: that they have a perfect analogy, and are both not only
composed of similar parts by their form, but also in their motions and
actions; as we have remarked in Chap. VI.
By the mixture of these two liquors I conceive the activity of the
organic molecules of each is stopped, and that the actions of one
counterbalance that of the other, insomuch that each particle ceasing
to move, remains in the place most analogous to itself, and that they
will naturally take the same position, and will dispose themselves in
the same order they held in the animal body; those that came from the
head will arrange themselves in the head of the foetus, those of the
back the same, and so of every other part; consequently they will form
a small organized being, in every thing like the animal from which they
are extracted.
It must be observed that this mixture of organic molecules of the two
sexes contains similar and different particles; the similar ones
are those which have been extracted from every part common to both
sexes. The different particles are those which have been extracted
from the parts whereby the sexes are distinguished; thus there is,
in this mixture, double the number of organic molecules to form the
head, or the heart, or such other parts common to both, whereas there
are only what are requisite to form the parts of the sex. Now the
similar particles may act upon each other without being disordered,
and collect together as if they had been extracted from the same body;
but the dissimilar parts cannot act on each other, nor unite together,
because they have not any relation; hence these particles will preserve
their nature without mixture, and will fix of themselves the first,
without the need of being penetrated by the others. Thus the molecules
proceeding from the sexual parts will be the first fixed, and all the
rest which are common to both, will afterwards fix indiscriminately,
whether they are those of the male or female, and form an organized
being which, in its sexual parts, will perfectly resemble its father,
if it is a male, and its mother if a female; but which may resemble
one another, or both, in all the other parts of the body.
It seems to me that if this was well understood, we shall in a great
measure be enabled to answer the objections made to the sentiments pf
Aristotle, and which might also be advanced against this system. The
question is, Why each individual, male and female, does not produce of
itself an animal of its own sex? It must be acknowledged this question
seems to carry weight with it; but having reflected a long time on this
subject I think I have found an answer, and which I shall endeavour to
explain.
It is certainly evident, from what we have said in the preceding
chapters, and the experiments we have described, that reproduction is
effected by the union of organic molecules returned from each part
of the body of the animal, or vegetable, into one or many common
reservoirs; and that they are the same molecules which serve for
nutriment and expansion of the body. This appears to me to have been
so clearly proved, that I apprehend no scruple can remain as to the
foundation of the theory; but I admit there may be some reason to ask,
Why each animal and vegetable does not produce its own likeness, since
each individual returns from every part of its body, and collects
in a common reservoir, all the organic molecules necessary for the
formation of a small organized being? Why then is not this organized
being formed? and why, in almost every animal, is a mixture of the
liquors of the two sexes required to produce an animal? If I content
myself with answering, that in almost all vegetables, and all kinds
of animals which multiply by cutting, that it appears the design of
Nature that each individual should increase its own species, and that
we must regard as an exception to this rule, the use which is made of
the sexes in other kind of animals; it may be said, that the exception
is more universal than the rule itself. This difficulty will be very
little weakened, if we were to say, that each individual perhaps would
produce its like, if it had proper organs, and contained the necessary
matter towards the nutriment of the embryo; because females have both
this matter, and organs, and yet do not produce either male or female
foetus without the intervention of the male; which intervention of
sexes in all animals is essential and absolutely necessary.
Although the testicles and seminal vesicles of a man, contain all the
necessary molecules to form a male, yet the local establishment and
arrangement of these molecules cannot be made, because the effect of
an union is prevented by the continual circulation of the seed both
by absorption, and the action of the new organic molecules which
constantly come into this reservoir from all parts of the body. The
same circumstances taking place with the organic molecules of the
female, is an evident reason why neither can produce of themselves,
because when the seminal liquors of the male and female are mixed,
they have more analogy to each other, than with the parts of the
body of the female where the mixture is performed. By admitting of
this explication, it may be asked, Why the common mode of generation
in animals does not agree with it; for, upon that supposition, each
individual would produce like snails, and impregnate each other, and
each individual receiving the organic molecules the other furnished,
the union would be made of itself, and by the sole power of the
affinity of these molecules among themselves? I own, if it was by this
cause alone the organic molecules could unite it would be natural
to conclude, that the shortest mode to perform the reproduction of
animals, would be to give to one individual both sexes. But it is
quite contrary to the general rule pursued by Nature, as this manner of
generation is confined to snails, and a small number of other animals.
This answer cannot be said to fully satisfy the question, as it merely
supposes the male does not produce, as it cannot receive any thing from
the female, and that having besides no proper viscera to contain and
nourish the foetus.
We may also suppose that the activity of the organic molecules, in
the semen of one individual, has need of being counterbalanced by the
activity or force of those of another individual, in order to fix
and bring them into a kind of equilibrium, a state of rest highly
necessary to the formation of the animal; and that this activity in
the organic molecules can only be counterbalanced by there being a
contrary action in those which come from the male, and those proceeding
from the female; so that, in this sense, all living or vegetating
beings must have two sexes, conjointly and separately, to produce
their resemblances. But this answer is too general to be entirely
clear; nevertheless, if we pay attention to all the phenomena, we
shall find some explanation resulting therefrom. The mixture of those
two liquors produces not only a male or female foetus, but also
other organized bodies, which have a kind of growth or expansion. The
placenta, membranes, &c. are produced at the same time as the foetus.
There are, therefore, in the seminal liquor of the male or female, or
in the mixture of both, not only organic molecules necessary for the
production of the foetus, but also those which form the placenta and
membranes. We know not from whence these molecules come, since there
is no part of the body, either of the male or female, from which they
could be sent back. From hence it seems it must be admitted, that the
molecules of the seminal liquors of each, being alike active, form
organized bodies every time they can fix, by acting mutually one on the
other: that the particles employed to form a male, will be those of the
masculine sex, which will fix the first, and form the sexual parts; and
that those common to both sexes will then fix indifferently to form the
rest of the body, and that the placenta and membranes are then formed
from the superabundant particles, which have not been used to form
the foetus; if, as we suppose, the foetus is a male, then there
remains to form the placenta, and membranes, all the organic particles
peculiar to the feminine sex which have not been employed; and also
all those of both which shall not have entered the composition of the
foetus, and which cannot be less than one half. So likewise, if the
foetus is a female, the same abundance will be left for the formation
of the placenta, and membranes, and the whole effects be the same,
excepting it will have the superfluity of the male, instead of that of
the female.
But, it may be said, that in that case the placenta and membranes
ought to become another foetus, which would be a female, if the
first was a male; and a male if the first was a female; for the first
having consumed the organic molecules of the sexual parts of only
one individual, and half those common to both, there remains all the
molecules of the sexual parts of the other individual, and the other
half of those common to both. To this I answer, that the first union
of the organic molecules prevents a second, at least, under a similar
form; that the foetus, being the first formed, exercises an external
power, which disorders the arrangement of the other organic molecules,
prevents the formation of a second foetus, and throws them into a
state from which the form of the placenta and membranes result.
We are assured by the experiments and observations we have made, that
every living being contains a great quantity of living and active
molecules. The life of the animal or vegetable appears to be only the
result of all the young lives (if that expression is permitted me) of
each of these active molecules, whose life is primitive, and appears
impossible to be destroyed. We have found these living molecules in
every living or vegetating being, and are assured, that they are alike
necessary for nutrition, and consequently, for the reproduction of
animals or vegetables. It is not, then, difficult to conceive, that a
certain number of those molecules united should compose a living being.
Each of these particles possessing animation, an assemblage of them
must be endowed with life, and thus these living organic molecules,
being common to all living beings, they necessarily form any particular
animal or vegetable, according as they are arranged. Now, this
arrangement absolutely depends on the form of the individuals which
furnish those molecules. If they are furnished by an animal, they will
arrange under the form of an individual like to it, exactly as they
were arranged when they served for the expansion of the animal itself;
but must we not then suppose that this arrangement cannot be made
either in animals or vegetables, but by the means of a kind of base,
round which the molecules might unite to form the foetus? Now, it is
plain, this basis is furnished by particles peculiar to the different
sexes, as I shall explain.
While the molecules of either sex remain by themselves, their action
produces no effect, because they are without any opposition from any
different kind of particles; but, when these molecules are mixed, then
there are dissimilar parts, and those serve for the base and point of
rest to the other molecules, and fix their activity.
In this supposition that the organic molecules, which, in the mixture
of the seminal liquors of the two individuals, represent the sexual
parts of the male, can alone serve for a base to the organic molecules
proceeding from every part of the female, and those peculiar to the
female sex as a base to them which are extracted from the male, we
might conclude, that the sexual part of the male infant is formed of
the organic molecules of the father, and from those of the mother, for
the rest of the body: and that, on the contrary, the female partakes
of its mother only in sex, and takes the rest of its body from its
father. Boys, therefore, ought, excepting the parts of the sex, to have
a greater resemblance to their mother than to their father, and girls
more to the father than to the mother; but this consequence is not,
perhaps, conformable to experience.
By considering, under this point of view, generation by sexes, we
should conclude it to be the most general mode of reproduction, as
it is in fact. Beings, whose organization is the most complete, as
animals, whose bodies compose a whole, which can neither be separated
nor divided, and whose powers are con-centered to one single point, can
only reproduce by this mode; because they contain only particles which
resemble each other, and whose union can only be made by different
particles, furnished by another individual. Those where organization is
less perfect, as that of vegetables, whose bodies may be divided and
separated without being destroyed, can be reproduced by other modes.
First, because they contain dissimilar particles; secondly, because
their forms not being so determinate and fixed as that of animals,
the particles may supply the functions of each other, and change
according to circumstances; as we see roots become branches, and shoot
out leaves when exposed to the air, which causes that the vegetable
particles obtain a local establishment, become fixed, and are enabled
to multiply, by various modes.
It will be the same with animals, whose organization is less perfect,
as the fresh water polypus, and others, which can reproduce by division
of their parts. These organized beings are not so much a single animal,
as a number united under one common covering, as trees are composed of
a multiplicity of young trees, (see Chap, II.) Pucerons, which engender
singly, also contain dissimilar particles, since, after producing
their young they change into flies which do not produce at all. Snails
communicate mutually these dissimilar particles, and afterwards they
both produce. Thus, in all known matters of generation, we see that the
requisite union of organic particles, can only be made by the mixture
of different particles, which serve as a basis capable of fixing their
motions.
If to the idea of the word _sex_, we give all the extent here supposed,
we shall say, that sexes are found throughout all nature; for then
sex will mean only the parts which furnish the organic particles,
different from the common particles, and which must serve as a fixed
point for their union. But, enough of reasoning on a question that
can be at once resolved, by saying, that God having created sexes, it
necessarily follows that animals should reproduce by their connection.
In fact, we are not made, as I have formerly said, to give a reason
for every _why_. We are not in a state of explaining _why_ Nature,
almost throughout her works, makes use of sexes for the reproduction of
animals, or why sexes exist; we ought, therefore, to content ourselves
with reasoning on what is, on things as they are, since we cannot go
beyond, by forming suppositions which will remove us from the sphere
we ought to contain ourselves in, and to which the small extent of our
knowledge is limited.
Quitting, therefore, all doubtful conjectures, I shall rest on facts
and observations. I find, that the reproduction of beings is formed in
many different manners; but, at the same time, I clearly perceive, that
it is by the union of the organic particles sent back from every part
of the individual, that the reproduction of vegetables and animals are
effected. I am certain of the existence of these organic and active
molecules in the seminal liquors of male and female animals and seed
of vegetables; and cannot doubt but every species of reproduction
is accomplished by the union of these organic molecules. Nor can I
doubt, that in the generation of animals, and particularly in that of
man, that the male and female particles mix in the formation of the
foetus, since we see infants which resemble both father and mother;
and what confirms this conclusion is, that those parts, common to both
sexes, mix promiscuously; whereas those never mix which represent the
sexual parts. For we every day see children with eyes like the father,
and the forehead and mouth like the mother; but we never find a like
mixture of the sexual parts; it never happens that they have the
testicles of the father, and the vagina of the mother, for even the
fact of hermaphrodites is very doubtful.
In the parts of generation of the two sexes in the human species,
there is so much resemblance, and so singular a conformity, that we
might be inclined to think those which appear so different externally,
are at bottom the same organs, only more or less developed; this was
the opinion of the ancients, and M. Daubenton's ideas on this subject
appear to me very ingenious.
The formation of the foetus is, then, made by the union of the
organic particles contained in the mixture of the seminal liquor
of both sexes; this union produces the local establishment of the
particles, which determines them to arrange themselves as they were in
the individuals which furnished them; insomuch, that the molecules,
which proceed from the head, cannot, by virtue of these laws, place
themselves in the legs, or any other part of the foetus. All these
molecules must be in motion when they unite, and in a motion which must
cause them to tend to a kind of centre, about which the union is made.
This centre, or fixed point, which is necessary to the union of the
molecules, and which, by its re-action and inertia, fixes the activity,
and destroys the motion, is, probably, the first assemblage of the
molecules which proceed from the sexual parts of the other individual;
they must arrange under the form of an organized body which will not be
another foetus, for the reasons we have before given.[AC]
[AC] In this, as in some other places, our author has gone into a
diffuse repetition which we have considered unnecessary and therefore
avoid.
On the whole, I conceive there are organic particles of the sexual
parts, which serve as a fixed point, or a centre of union, around which
all the other parts that form the embryo collect. I speak of it only
as probable; but as they are the only particles which differ, I have
thought it more natural to imagine, that it is around these different
particles the union is formed than those which are common to both sexes.
We have before observed, that those who have imagined the heart was the
first formed, are deceived: those who say it is the blood, are no less
so. All is formed at the same time. If we only consult observation, the
chicken is seen in the egg before it has been sat upon; we perceive the
spine of the back and the head, and, at the same time, the appendages
which form the placenta. I have opened a great number of eggs, before
and after incubation; and I am convinced, by my sight, that the chicken
exists entirely in the middle of the cicatrice, the moment it comes
from the body of the hen. The heat, communicated to it by incubation,
only expands the parts by setting the liquors in motion; but it is not
possible to determine which parts of the foetus are fixed in the
instant of formation.
I have always said, that the organic molecules were fixed, and that
their uniting was caused only by their loss of motion. This appears
to me certain: for, if we separately examine the seminal liquor of
the male and female, we shall see an infinity of small bodies in
great motion, but being mixed, their motion is instantly suspended,
and heat is necessary to renew their activity; for the chicken which
exists in the centre of the cicatrice is without any motion before
incubation; and even twenty-four hours after, when it begins to become
perceptible with a microscope, there is not the least appearance of
motion then, nor even the day following. During the first day it is
only a small white mucilaginous mass, which is of a consistence on
the second, and insensibly increases, but whose motion is very slow,
and does not at all resemble that of the organic particles which move
rapidly in the seminal liquor. Besides, I have reason to say, that
this motion of the organic molecules is absolutely destroyed; for if
we keep an egg without exposing it to a degree of heat necessary to
expand the chicken, the embryo, although formed entirely, will remain
without any motion; and the organic molecules of which it is composed,
will remain fixed without being able to give motion and life to the
embryo which has been formed by their union. Thus, after the motion
of the organic molecules has been destroyed, after the union of these
molecules, necessary to form an animal body, there is still an external
agent required to animate and give it life and motion; and this agent
is heat, which, by rarefying the liquors, obliges them to circulate
and put also every organ in action, which afterwards do no more than
develope and grow, provided that this external heat continues to assist
them in their functions.
Before the action of this external heat, not the least appearance of
blood is to be seen; and it is not till twenty-four hours after, that
I have perceived any change in the colour of the vessels. The blood
first appears in the placenta, which communicates with the body of
the chicken: but this blood seems to lose its colour as it approaches
the body of the animal; for the chicken is entirely white, and we
with difficulty discover in the first, second, and third days after
incubation, a few small sanguinary points which are close to the body
of the animal, but which seem not to make part of it, although it is
these sanguinary points which afterwards form the heart. Thus, the
formation of the blood is a change occasioned in the liquors by the
motion heat communicates to them, and this blood is formed even out of
the body of the animal, the whole substance of which is then only a
kind of mucilage, or thick jelly.
The foetus, as well as the placenta, derives the necessary nutriment
for expansion, by a kind of absorption, and they assimilate the organic
parts of the liquor in which they float: for the placenta cannot be
said to nourish the animal, no more than the animal nourishes the
placenta; since, if the one nourished the other, the first would soon
appear to diminish, while the other increased, whereas both increase
together, I have indeed observed in eggs, that the placenta at first
increases much more in proportion than the foetus, and therefore
it may nourish the animal, or rather convey the nutriment to it, by
intussusception.
What we have just said concerning the chicken, is easily applied
to the human foetus, which is formed by the union of the organic
molecules of the two sexes. The membranes, and placenta, are formed
from the superabundance of the particles which have entered into the
composition of the embryo: which is then inclosed in a double membrane,
where there is also a quantity of liquor, which is, perhaps, at first,
but a portion of the semen of the father and the mother; and as the
foetus does not quit the matrix, it enjoys, from the instant even
of its formation, an external heat necessary for its expansion; this
heat communicates a motion to liquors, and sets the organs in play, and
blood is formed in the placenta, and in the body of the embryo, by the
motion occasioned by this heat. It may be even said, that the formation
of the blood of the infant is as independent of the mother, as that
which passes into the egg, is of the hen which hatches it, or of the
oven which heats it.
It is certain, that the foetus, placenta, and membranes, grow by
intussusception: for, in the earliest days of conception, the pouch,
which contains the whole product of generation, is not adherent to
the matrix. De Graaf, in his experiments on doe rabbits, made these
globules, wherein the whole business of generation lies, move about
in the matrix. Thus, in the first stages, they increase and grow by
drawing nutriment from the liquors which bathe the matrix, to which
they are afterwards attached by a mucilage, in which small vessels are
formed with time, as we shall hereafter explain.
But, not to quit the subject, let us return to the immediate formation
of the foetus, on which there are many remarks to be made, both as
to its situation, and the different circumstances which may prevent or
stop its formation.
In the human species, the seed of the male enters into the matrix, the
cavity of which is considerable; and when it meets with a sufficient
quantity of female semen, a mixture of the organic particles succeed,
and the formation of the foetus ensues: the whole, perhaps, is done
instantaneously, especially if the liquors are both in an active and
flourishing state. The place where the foetus is formed, is the
cavity of the matrix, because the seed of the male can enter there
more easily than into the trunks; and as this viscera has but one
small orifice, which is always shut, excepting when the ardour of
love causes it to open, the materials of generation remain there
with safety, and scarcely ever reissue but by rare and unfrequent
circumstances: but as the liquor of the male sprinkles the vagina,
before it penetrates the matrix, by the activity of the organic
molecules which compose it, it may go farther into the trunks, and,
perhaps, into the ovarium. As the liquor of the female has already its
perfection in the glandular bodies of the testicles, from which it
flows and moistens the trunks and other parts before it descends into
the matrix, and as it may issue out of the vacuities left around the
neck of the matrix, it is not impossible, that the mixture of the two
liquors may be made in all these different places. It is, therefore,
probable that foetuses are often formed in the vagina, but which fall
out as soon as they are formed, because there is nothing to retain
them. It may also sometimes happen, that foetuses are formed in the
trunks; but this case is very rare, and cannot happen but when the
seminal liquor of the male enters the matrix in great plenty.