Denmark, Sweden, and Finland, and one of them has also been sent from
Oran. The genus is small, and may have been overlooked in other
countries, although its appearance is sufficiently distinct and marked
to have caught the eye. It is as lithe and active as a Malay, as black
as a negro, and as hairy as a gorilla, looking like a little ursine
sweep.
The genus EUCERA, of which we have but one representative, although
considerably more than fifty species are known, has not so wide a range
as might be expected from their numbers. Our own is found throughout
Europe and in Algeria. Other species occur in Russia, the Morea,
Albania, Dalmatia, and Egypt. In Asia some are found in Syria, and at
Bagdad; and from the New World they have been sent from Cayenne and the
United States.
The genus ANTHOPHORA, to which the genus _Saropoda_ is very closely
allied,—so closely, indeed, that by the celebrated hymenopterologist Le
Pelletier de St. Fargeau the species of both are incorporated
together,—has, even as now restricted, a world-wide dissemination, and
numbers nearly a hundred and fifty species. Several of our own occur
throughout France and Italy and the whole of northern Europe, and even
among the Esquimaux in the arctic regions, showing that a bridal bouquet
may be gathered even there; for where bees are flowers must abound.
The genus in other species shows itself in the south of Europe, viz. in
Spain, Sicily, the Morea, and Dalmatia; by way of Syria and Arabia Felix
it passes down to Egypt and occurs in Nubia and also in Algeria. It dots
the western coast of Africa at Senegal and Guinea, and has been
discovered in the Canaries, and again makes its appearance at the Cape
of Good Hope, rounding it to Natal. It travels round the peninsula of
India, being found at Bombay, in Bengal, and in the island of Ceylon,
and passes onward by way of Hongkong to northern China, where, dipping
to the Philippines, it next occurs in Australia. In the New World it is
found on its western side at Chili, and traverses that continent to
Paraguay and Pará, and has been sent from the West India Islands of
Cuba, St. Domingo, and Guadaloupe. From Mexico, where we next find it,
it passes to Indiana, and occurs throughout the United States, and thus
completes its progress round the world. About one hundred and thirty are
known.
The genus SAROPODA is closely allied to _Anthophora_, as closely as
_Heriades_ is to _Chelostoma_, and is very limited in numbers, ten only
being known, and but one of which is native with us. The genus occurs
throughout France and Germany, and has been sent from Russia, Egypt,
South Africa, and Australia, thus having a very wide range
notwithstanding the paucity of its species.
The very pretty genus CERATINA, although numbering but few
species,—fewer than thirty,—and although not found in Australasia, is
widely scattered throughout the Old and the New Worlds. Our own species
inhabits as far north as Russia. Other species occur throughout France,
and in the south of Europe, and show themselves in the Morea, and in
Albania. North, South, and Western Africa possess the genus, it being
found in Algeria and at the Cape of Good Hope, and in the intervening
district of Senegal. It has been brought from Ceylon and Bengal, and
also from the north of India. It reaches China by way of Java and
Hongkong: and in the New World has been found in the Brazils and
Cayenne, in the Southern, and throughout the United States in the
Northern continent.
The genus NOMADA is the first of the genuine parasitical bees, and about
the habits of which no doubt can be entertained; certainly not the same
as attaches both to _Hylæus_ and _Sphecodes_, among the _Andrenidæ_. The
parasitical habits of Nomada are evident and unmistakable. This is the
handsomest genus, in variety of colour and elegance of form, of all our
native bees, but the species are never conspicuous for size. They have
much of the appearance of wasps, and are often mistaken for them even by
entomologists, who have not paid attention to bees. Many of our native
species seem limited to our own islands: others of our species occur in
France and Germany, and through Denmark in direct line to Lapland,
turning down into Russia, and have been caught as far south as Albania.
One of our species, or so like as to want distinguishing
characteristics, is found in Canada. Did ours migrate there? and how?
The genus is of wide distribution, but occurs only north of the Equator,
where it spreads from Portugal to the Philippine Islands. It is found in
Siberia and Northern China, whence through the Philippines it passes to
Tranquebar, then up to Northern India, and thence by Bagdad to the Morea
and Albania, and dips down to Northern Africa at Tunis, and on to Oran
and Tangiers, and completes its circuit in Portugal. It is doubtless
parasitical upon many more genera and species than we find it infest in
this country, although all that the several species pair off with here
are not fully designated, especially among the _Andrenæ_, and smaller
_Halicti_. The number of species, British and foreign, known to
collectors approximate to a hundred.
The genus MELECTA is another handsome parasitical insect. This is always
a dark beauty, and is very limited in species, for, as far as they may
be estimated from the contents of collections, its numbers do not reach
twenty. Our own species occur throughout the whole of Europe, north and
south. Others are found in Sicily, Albania, the Morea, and show
themselves at Bagdad. The genus has been sent from the Canaries, and
crosses the tropics into Chili, but does not seem to have occurred
elsewhere in either North or South America, although one of the genera
(_Eucera_) on which, with us, it is parasitical, is found in the latter
country, and the other genus (_Anthophora_), which it also infests, is
found throughout the world, excepting in Australasia. In all those
countries, the closely-allied exotic genus _Crocisa_, which is very
numerous in species, may supply its place.
The elegant genus EPEOLUS occurs in our own species throughout northern
Europe, as high as Lapland, and is found also at the southern extremity
of the continent of the Old World, at the Cape of Good Hope. It has been
brought from Sicily, and other species come from Siberia. The genus in
America passes down from the United States, by way of Mexico, to the
Brazils, where it crosses the southern continent, having been
transmitted from Chili. It is very limited in the number of its species,
considering its wide diffusion, for not more than twenty are registered.
It is almost identical in distribution with the genus _Colletes_, upon
which it is with us parasitical. The species are never so large as those
of the preceding genus, _Melecta_.
The genus STELIS is limited both in number of species and distribution,
although the spots whence it has come are wide apart. Our own species
are found throughout France and northern Europe, as far as Finland.
Other species occur in North America, and the Brazils, but the whole
number yet described is under ten.
The remarkable form in both sexes of the genus CÆLIOXYS occurs in
identity with our own species throughout France and Austria, and spreads
north to Finland and Russia, and through all the intervening countries.
It is singular that it should not be recorded from southern or
south-western Europe, as it is found in Oran. Other species of the genus
have been found in northern Africa, Egypt, and Algeria. On the western
coast of Africa it has been caught on the Gambia, at Sierra Leone, and
on the coast of Guinea. It doubles the Cape of Good Hope, where it is
found extending its range to Port Natal. From Asia we have it from
Turkey, and again from India. It has been sent from the hither side of
South America, from the Brazils, and separately from Pará, and occurs at
Cayenne, and in the West India Islands, Cuba, and St. Thomas’s, and
extends as high in North America, through the United States, as Canada.
It is quite probable that it has as wide a range as the bees upon which
it is parasitical (_Megachile_), although it has not yet come from such
extensively-spread localities. More than fifty species are known, but
some of our own have not yet been enumerated amongst those found
elsewhere.
The genus MEGACHILE, which embraces the most renowned of the mechanical
bees, is extremely cosmopolitan, spreading north and south, east and
west; and is also very abundant in the numbers of its species, the
census extending to not far short of two hundred. Some one, or several
of our species, although other species are limited to our own
country,—spread through Italy and France, and all the countries of
northern Europe to the high latitude of Lapland, which is higher than
where even one of ours (viz. the _M. centuncularis_) is again found,
which occurs in Canada and at Hudson’s Bay. The genus also frequents
southern Europe, in Spain, Sicily, and Albania, and in the East, in the
Caucasus and Dalmatia. It traverses Turkey by Bagdad to India, having
been captured in Nepaul, and it descends southward in the Indian
peninsula, where it has been found at Bombay. From India it stretches to
the Mauritius, thence across the Indian Ocean to Java, and thence to
Hongkong and northern China. It then dips to the Philippines, and
doubtless through the islands of the Indian Archipelago to Australasia,
from which continent none are registered from its northern and eastern
settlements, but species abound along its southern edge from Western
Australia, through Adelaide to Tasmania. The genus has been brought from
the West India islands, St. Thomas’s, St. Croix, and Cuba: it is found
upon the main from Mexico, descending to the Brazils. It skirts all the
coasts of Africa, being discovered in Egypt and Algeria, along the
western coast by the Gambia, Senegal and Sierra Leone to Guinea, and the
island of Fernando Po, and then again occurs at the Cape of Good Hope.
Ascending the eastern coast by Natal, it stretches to Abyssinia. The
species are very abundant in India, Africa, and Australasia.
The genus ANTHIDIUM, although very numerous in species, and differing
more remarkably in form amongst themselves than most other genera, has a
far less extensive range, no species having been found in Australasia or
India, although it occurs in Arabia, Syria, and Mesopotamia. Our own
solitary species occurs in France, Italy, and the whole of northern
Europe, extending to Finland. In southern Europe the genus inhabits
Sicily, Spain, the Morea, Albania, and Dalmatia, and is also very
abundant in Southern Russia. In Africa it is found in Nubia and Algeria,
and on its north-western edge in Barbary, whence it descends by the
Gambia and Sierra Leone to the Cape of Good Hope, and thence reaches to
Natal. It is then found in Chili, and crossing the South American
continent occurs in the Brazils, whence it ascends to Cayenne, and, by
way of Mexico, to the United States. The number of species recorded
exceed a hundred.
The remarkable genus CHELOSTOMA is very limited in the numbers of its
species, of which less than a dozen are known; as also in the extent of
their distribution. Our own are found throughout northern Europe, as far
as Lapland, and in Russia. In southern Europe they occur in the Morea,
and the genus has been discovered in Georgia in North America.
The closely-allied genus HERIADES seems limited to a European
habitation, and occurs only in our own solitary species, but it ranges,
like the preceding, to the high latitudes of Lapland.
ANTHOCOPA seems limited to our own country and France, possibly only
from its having been associated from similarity of general habit with
the genus _Osmia_. Only one species appears to be known, but this has a
world-wide celebrity, from the interesting account given by Réaumur, of
its hanging its abode with symmetrical cuttings from the petals of the
poppy.
The genus OSMIA, although not including such able artisans as
_Megachile_, still has in its species very constructive propensities.
Indeed, all the bees which convey the pollen on the under side of the
abdomen, are more or less builders or upholsterers. The genus has a wide
range, and is tolerably numerous, numbering more than fifty species.
Some of our own occur throughout Europe, and, like the two preceding
genera, are found in the highest continental latitudes. Some of ours
also occur in Algeria and the Canaries, other species in Albania and
Moravia. In Africa they are found in Egypt, Barbary, and Port Natal, and
in the New World from Florida, in the United States, through Nova Scotia
to Hudson’s Bay.
The genus APATHUS, which is parasitical upon _Bombus_, and to the
uninitiated has all the appearance of this genus, seems to be the only
instance of a parasitical genus of bees so closely resembling the
_οἶτος_, (as we may, perhaps, for the sake of avoiding a periphrasis, be
allowed to call the bee upon which the parasite is found,) as to be so
easily liable to be mistaken for it, and which was indeed the case by
even such a sagacious entomologist as the distinguished Latreille; but
Kirby had already noticed the difference, suggesting its separation from
_Bombus_, until about the time that St. Fargeau was induced to propose a
distribution of the Hymenoptera, based generally upon economy and
habits, to which he had been led by a refining investigation of
structure, that the distinguishing difference was appreciated, and used
generically, by Mr. Newman. This difference, like many other simple
facts, now that it has been found, is very obvious. It consists in the
genus having no neuters, and the female of the species no polliniferous
organs, but the determination of the legitimate males, by means other
than empirical, is still difficult. In our own species this genus ranges
throughout northern Europe, as high as Lapland; a cause for which we
shall discover when we trace the geography of the next genus, _Bombus_.
One species different from any of ours occurs in the Brazils, and others
are found in the Polish Ukraine, and in the United States of North
America. The genus appears extremely limited in numbers, for although
nearly a hundred of the genus _Bombus_ are known, _Apathus_, in
collections, seems limited to ten. This may perhaps arise from want of
due observation or from the neglect of their careful separation from
that genus, but our own species are far from co-extensive with our
native species of _Bombus_.
The genus BOMBUS, although with some southern irrepressible
propensities, it being found within the tropics in a few instances, is
essentially a northern form, which is strongly indicated in its downy
habiliments, for it is clothed in fur like the Czar in his costly
blue-fox mantle. In the Old World its range extends to Lapland, whither
it is followed, as previously noticed, by its parasite _Apathus_, and in
the New World to Greenland, where one species seems an autochthon,
perhaps originating there when the land was still verdant, and grew
grapes, long before the age of Madoc. Other species occur far away to
the north of east, booming through the desolate wilds of Kamtchatka,
having been found at Sitka; and their cheerful hum is heard within the
Arctic circle, as high as Boothia Felix, thus more northerly than the
seventieth parallel. They may, perhaps, with their music often convey to
the broken-hearted and lonely exile in Siberia, the momentarily cheering
reminiscence of joyful youth, and by this bright and brief interruption
break the monotonous and painful dullness of his existence, recalling
the happier days of yore: but the flowers of humanity, here typified by
the natural flowers which attract these stray comforters, will one day
spring where the salt of tears now desolates, and thus the merry bees
have sweetness for even these poor outcasts, and froth their bitter cup
with bubbling hope.
In the south of Europe the genus occurs in Austria, the island of Zante,
and the Pyrenees. It is found in Syria, the island of Java, in China at
Chusan and Silhet, and also in northern India; and, although crossing
the tropics to fix itself at Monte Video, at the mouth of Rio de la
Plata, in Africa it appears to be found at Oran only; nor does it occur
in Australasia. In South America it is also found at Pará and Cayenne,
and on the opposite side at Columbia, Quito, and Chili, and passes up
the isthmus to California, and thence to Mexico, whence it extends to
the island of Antigua.
-------
The genus APIS, or the HIVE BEE,—which perhaps in its past and present
utility to man, may successfully compete in the aggregate with the
silkworm,—with true regal dignity comes the last of the series of
genera. The whole array of her precursors, who marshal her way, and
derive their significance and importance from the more or less direct
resemblance in structure and function to her, deduce their common name
of “Bees” from this relationship, and consequently from her. Long before
their existence had been traced by the observer of nature or by the
naturalist, the comb of the BEE had dropped in exuberant luxuriance its
golden stores for the gratification of mankind. This little creature had
garnered, from sources inaccessible to man, the luscious nectar
concealed within the bosom of the flower, whose exquisitely beautiful
varieties, in form, colour, and fragrance, had delighted his sight and
his smell long before he had been led by accident to discover that these
industrious little workers collected into their treasury, from those
same flowers, as exquisite a luxury for his taste, as they themselves
had yielded to his other senses. Thus the earliest records speak of
honey, and of bees, and of wax; and the land of promise to the restored
Israelites, was to be a land flowing with milk and honey.
Réaumur, whose observations upon bees had been pursued with such patient
and indefatigable perseverance, combined with such minute accuracy, and
then recorded so agreeably, and who conceived the possibility of
establishing a standard of length, for the common use of all nations, to
be derived from the length of a certain number of the honey-cells of the
comb, to which notion he was doubtless led by their mathematical
precision and uniform exactitude, appears to have been unaware of the
existence of other species of the genus, and hence he assumed, in his
ignorance of this fact, that in all countries they were alike.
Travellers had, even for more than a century before, mentioned different
kinds of honey, derived from different kinds of bees, which, however,
Réaumur does not, from this circumstance, seem to have known. Had he
been acquainted with it, his philosophical accuracy of observation and
habit of reflection would certainly have assumed the possibility of
differences of size in the cells of the different bees, and he would
have waited until opportunity had given him the power of determining
whether this mode of admeasurement could be safely adopted as certainly
being of universal prevalence. It is to be wondered at also, that he did
not weigh the possibility that climatic differences in the distribution
of even the _Apis_ _mellifica_ might have involved discrepancies, by the
effects constantly seen to be produced by climate, and which would have
shown that the standard which he sought to establish could not be relied
on.
Collections exhibit about sixteen species of the genus _Apis_, whose
natural occurrence is restricted to the Old World, for although the
genus, especially in the species _A. mellifica_, has been naturalized in
America, and also in Australasia, and in some of the Islands of the
Pacific, these were originally conveyed thither by Europeans. Those
countries possess representatives of the genus with analogous attributes
and functions, in two other genera, which fulfil the same uses. It is
remarkable that the Red Indians used to note the gradual absorption of
their territory by the White Man, through the forward advance of his
herald _Apis mellifica_. This species has also been carried to India, to
the Isle of Timor, and to northern, western, and southern Africa, in all
which countries it is thoroughly naturalized, although they all possess
indigenous species, which are quite as, or perhaps more largely,
tributary to their inhabitants. Observation has not hitherto confirmed
the identity of the manners of these exotic species with our own, owing
to the deficiency of observers with the enthusiasm requisite to follow
their peculiarities with the patience of a Réaumur, a Bonnet, or a
Huber. That they are quite or all but similar, exclusively of
differences of size, both in their habits and their nests, may be
inferred from their identity of structure. We know that they consist of
three kinds of individuals—neuters, females, and males,—and that their
combs are made in cakes built vertically, formed of hexagonal contiguous
cells, which are placed bottom to bottom, and overlap each other in the
same strengthening position as do ours; and also that the cells wherein
the males are developed are oval, larger than the honey-cells, and less
uniform. With all these similitudes it is fair to suppose that their
economy may be the same; but their honey-cells, from their smaller size,
(the bee which produces them being smaller,) have a more elegant
appearance; and it is concluded from the largeness of the nest, taken
conjunctively with the smallness of the cells, and of the bees
constructing it, that the communities thus associated must in their
collective number be considerably larger than those of our hives.
Instinct, as expressed in the habits, is as sure a line of separation,
or means of combination, as structure, and is corroborative in tending
to preserve generic conjunction in its inviolability. And, conversely,
with certainty, is indicated that such-and-such a form, in the broad and
most distinguishing features of its economy, is essentially the same in
every climate. The habits, therefore, in whatever country the genus may
occur, may be as surely affirmed of the species, from the knowledge we
have of those at home, as if observation had industriously tracked them.
This is especially the case in a genus, the species of which present
such a peculiar identity of _structure_ as does _Apis_, whose specific
differences are derived only from colour and size, and this identity is
a peculiarity, so far as I have observed, rarely found in other genera,
numbering even no more species, but wherein slight differences of
structure often yield a subsidiary specific character, complete
structural identity being almost solely incidental to the genus _Apis_.
The importance of honey and wax throughout the world, as well for the
ceremonies of religion, as for the service of the arts, and for medical
or domestic purposes, is attested by the vigilance, care, and assiduity
with which bees are tended in every country. Although sugar, since its
introduction to those northern countries which have not been favoured by
nature with the cane that yields it, has superseded for ordinary uses
the produce of the hive, this still continues serviceable for many
purposes to which sugar cannot be applied. It is used in many ways in
pharmacy, and still retains in the interior of some continents, owing to
the deficiency of sugar, arising from the difficulties and expenses of
transit, all its primitive uses. In the East, even in countries
producing sugar in abundance, honey is extensively employed for the
preservation of fruits, which in their ripe state in those hot climates
would rapidly lose their fulness of flavour were they not thus
protected,—honey here being esteemed superior to sugar in the
circumstance of its not crystallizing by reason of the heat, and also
from its applicability to this use in its natural state.
This is especially the case in China, where a conserve of green ginger,
and of a fragrant orange (the _Cum Quat_), are in high repute, and which
are peculiarly grateful to Europeans on the spot. These, however, are so
delicately susceptible of change of climate, that they lose some of the
aroma that constitutes much of their attraction, upon transportation,
and, indeed, like many kinds of Southern wines, can be appreciated only
within their own country, from their extreme delicacy and tendency to
spoil.
Honey is a very favourite food and medicine with the Bedouins in
Northern Arabia. Bees make their hives in all the crevices of rocks in
Hedscha, finding everywhere aromatic plants and flowers. At Taif, bees
yield most excellent honey, and the honey at Mecca is exquisite. At
Veit-el-Fakeh, wax from the mountainous country of Yemen is exchanged
for European goods and for spices from the further Indies. In Syria and
Palestine we find bees abound. At Ladakiah there are large exports both
of honey and wax; and the honey of Ainnete, on the declivities of the
Lebanon, is considered the finest of the whole of that mountain-range.
Antonine the Martyr, in the seventh century, speaks of the honey of
Nazareth being most excellent, and in the present day bees are
extensively cultivated at Bethlehem, for the sake of the profit derived
from the wax tapers supplied to the pilgrims. Some of the members of the
German colony at Wadi Urtas speak of the purchase of eleven beehives at
this place, and express themselves as very sanguine of an abundant
harvest from the luxuriance and profusion of flowers, although they say
the bees are smaller than those of Westphalia, and are of a
yellowish-brown colour. The eastern side of this peninsula, especially
the district of Oman, is wholly destitute of bees, contrasting thus
unfavourably with its western fertility.
The enormous quantities of honey produced may be comparatively estimated
by the collateral production of beeswax, which it exceeds by at least
ten to one. When we reflect upon what masses of the latter are consumed
in the rites of the Roman Catholic and Greek churches throughout the
many and large countries where those religions prevail, we shall be able
to form a general estimate of the extensiveness and universality of the
cultivation of bees. Nor are those the only uses to which wax is
applied, and the collective computation of its consumption will show
that bees abound in numbers almost transcending belief.
The name of _bougie_ for wax-candle or taper, is used by all the
languages of the south of Europe, and is derived from the name of Bugia,
a town of Northern Africa, whence, even as long back as the time of the
Roman Empire, wax was obtained to make candles for lighting. The
inhabitants of Trebizonde paid their tribute to the Roman Empire in wax.
Both honey and wax are largely employed in pharmacy, and were also, in
ancient times, both extensively used in embalming. The honey of Mount
Hymetta in Attica, and of Hybla in Sicily, were each in as high repute
in classical countries as is that of Narbonne in Languedoc, by reason of
its choice delicacy, with us, and throughout France. Distributed over
the wide pastures of the Ukraine, every peasant has his store of hives,
which frequently, in their harvests, realize more largely than their
crops of grain,—multitudes of that peasantry computing as important
items in the estimate of their wealth the number of their beehives,
which often exceed five hundred to the individual possessor. In Spain
and Italy bees are largely cultivated; and in the former country many a
poor parish priest, the religious monitor of an obscure hamlet, can
count his five thousand.
In countries so rich in the productions of Flora, whose seasons there
are perennial, and which fluctuate only in special locality, bees are
removed to and fro to meet these peculiarities. Thus in the south of
France, where large tracts are cultivated with aromatic shrubs and
flowers, for the distillation of essential oils and fragrant waters, the
hives of bees are moved up and down the adjacent rivers upon rafts, as
the flowering of the crops succeed each other. In Italy, Spain, and
Southern Russia, the same practices are pursued, although we have no
detailed accounts of the precise spots; but we know from Niebuhr,
Savigny, and Sir Gardiner Wilkinson, that upon the Nile it is customary
thus to transport the bees from flower-region to flower-region upon
rafts containing about four thousand hives, each numbered by the
proprietors of the hives for identification, who thus double the seasons
by continually shifting their bees from Lower Egypt to the Upper Nile
and back again.
In ancient Greece also, they were conveyed for this purpose from Achaia
to Attica; in the former of these provinces, owing to its higher
temperature, flowers had passed their bloom before spring had opened in
the latter. All these circumstances tend to show that the experience of
bee-masters, both ancient and modern, has ascertained that their insects
have not a very extensive range of flight.
Of the fact that the honey of bees is not always salutary to man, there
is a remarkable instance recorded in Xenophon, in his narrative of the
retreat of “The Ten Thousand,” who reports that upon falling in with
quantities of it, in Asia Minor, those who indulged in its enjoyment
were seized with vertigo, or headache, and violent diarrhœa, attended
with sickness, but which had no fatal consequences, although they did
not recover from its injurious effects for a couple of days, and were
left then in a very prostrated condition. The celebrated physician and
botanist Tournefort, when travelling in the East, towards the end of the
seventeenth century, found, in the neighbourhood of Trebizonde, an
excessive luxuriance of the flowers of the _Rhododendron ponticum_ and
of the _Azalea pontica_, which, although sumptuous in their blossoms,
were held in bad repute by the inhabitants, who ascribed to their odour
the deleterious effect of causing headache and vertigo. He was thence
induced to surmise that these had possibly been the flowers the bees had
extracted the honey from which had been so baneful to the troops of
Xenophon.
But it seems that bees themselves cannot collect with impunity the honey
of noxious flowers, for they are occasionally subject to a disease
resembling vertigo, from which they do not recover, and which is
attributed to the poisonous nature of the flowers they have been
recently visiting.
Several different kinds of honey and wax have been described, but some
degree of uncertainty exists as to whether they are all the produce of
genuine species of the genus _Apis_; for it will be found, in a rapid
notice I purpose giving of the more conspicuous genera of foreign bees,
that there are two exotic genera of this section of the family, both
social in their habits, and which both produce the same materials; there
is a wasp also that makes honey. But of all the many kinds of honey
noticed, the green kind furnished to Western India by the island of
Réunion, the produce of an _Apis_ indigenous to Madagascar, but which
has been naturalized in the French island, and also in the Mauritius, is
perhaps the most remarkable. It is of a thick syrupy consistency, and
has a peculiar aroma. It is much esteemed upon the most proximate coasts
of the peninsula of India, where it bears a high price. Whether its
greenness of colour is derived from the flowers which this species
frequents, or whether it be incidental to the nature of the bee, has not
been ascertained, but the honey of the South American wasp, the sole
species producing the material, has also a green tinge.
Nature has assigned the task of thus catering for man, by collecting and
garnering from the recondite crypts within the blossoms of flowers, to
about sixteen species congenerical with our honey-bee, but sufficiently
differing. As I have before noticed, the species of this genus greatly
more resemble each other in structure than perhaps do the species
collocated within any other genus of insects, and whence may be inferred
an exact similitude of habits, although as yet unconfirmed by direct
observation.
The second European species, the _Apis Ligustica_, or Ligurian bee, is
rather larger, but very like ours, and inhabits the whole of the north
of Italy, its occupation of that country extending from Genoa to the
vicinity of Trieste; its progress further north being impeded by the
Alps of Switzerland and the Tyrol. It is also found in Naples, and may
likewise spread to the Morea, Turkey, and the Archipelago of Greece, and
is perhaps the bee noticed by Virgil. Either this species, or possibly
one distinct from ours, is that which is so extensively cultivated in
Spain, although ours is found in Barbary.
Another smaller kind, the _Apis fasciata_, has been cultivated in Egypt
from time immemorial, and which yielded its abundant harvests for the
gratification of the ancient Romans. Only five other distinct species,
so far as is yet known to us, appear to occupy the vast continent of
Africa,—two on its western coast at Senegal and Congo, the _A.
Adansonii_ and the _A. Nigritarium_; two in Caffraria, the _A.
scutellata_ and the _Apis Caffra_. That at Madagascar, and doubtless on
the adjacent mainland, which has also been naturalized in the Mauritius
and at Réunion, is the _Apis unicolor_, which produces the green honey
mentioned above.
India, however, at present appears to be the true metropolis of the
genus. Further discoveries in Africa may hereafter give that vastly
larger continent the predominancy; but there is no doubt that, so far as
present information extends, India has the superiority. Thus _Apis
dorsata_, _Apis nigripennis_, and _Apis socialis_, are cultivated in
Bengal, the latter being also found along the Malabar coast and at Java.
It is singular that the only instance of the occurrence of the very
distinct genera of _Apis_ and _Mellipona_, both honey-storing genera,
yet known to exist indigenously in the same locality, is found in this
island. At Pondicherry and its vicinity are found _Apis Delessertii_ and
_Apis Indica_. This latter bee is extensively cultivated, and its hives
are perhaps the most largely inhabited of any of the species; the
numbers occupying a single nest being estimated at above eighty
thousand.
From India also, but to which no special locality is assigned, come
_Apis Perrottetii_, _Apis lobata_, as likewise _Apis Peronii_, which is
equally native to the Isle of Timor. The honey produced by this last bee
is yellow, more liquid than ours, and of a very agreeable flavour.
Thus science dissipates the popular supposition, that a multiplicity of
the individuals of one species of this insect produces the tons of wax
and the myriads of gallons of honey that are annually consumed.
Which of these bees first benefited the human race, in its primitive
seat, and before the multiplication of mankind forced them to take
divergent courses from the cradle of their birthrace, “to people the
whole earth,” it is impossible to say. And it is equally impossible to
conjecture whether, like man, they by this course of migration have
assumed the features they now exhibit of distinctly different species;
yet they do not vary so considerably among themselves as do many other
creatures that have come under the direct influence of man,—the chief
differences consisting in the comparatively slight distinctions of
colour and of size, but which are sufficiently marked to constitute them
good species.
The earliest manuscript extant, which is the Medical papyrus, now in the
Royal Collection at Berlin, and of which Brugsch[2] has given a
facsimile and a translation, dates from the nineteenth or twentieth
Egyptian dynasty, accordingly from the reign of Ramses II., and thus
goes back to the fourteenth century before our era. But a portion of
this papyrus indicates a much higher antiquity, extending as far back as
the period of the sovereigns who built the Pyramids, consequently to the
very earliest period of the history of the world.
Footnote 2:
‘Recueil de Monuments Égyptiens dessinés sur les lieux.’ In Three
Parts. 4to. Leipzig, 1862.
It was one of the medical treatises contained within the Temple of Ptah,
at Memphis, and which the Egyptian physicians were required to use in
the practice of their profession, and if they neglected such use, they
became responsible for the death of such patients who succumbed under
their treatment, it being attributed to their contravening the sacred
prescriptions. This pharmacopœia enumerates amongst its many
ingredients, honey, wine, and milk; we have thus extremely early
positive evidence of the cultivation of bees. That they had been
domesticated for use in those remote times, is further shown by the fact
mentioned by Sir Gardiner Wilkinson of a hive being represented upon an
ancient tomb at Thebes.
It may have been in consequence of some traditional knowledge of the
ancient medical practice of the Egyptians, that Mahomet, in his Koran,
prescribes honey as a medicine. One of the Suras, or chapters, of that
work, is entitled ‘The Bee,’ and in which Mahomet says:—“The Lord spake
by inspiration unto the Bee, saying, ‘Provide thee houses in the
mountains and in the trees [clearly signifying the cavities in rocks and
hollows of trees, wherein the bees construct their combs], and of those
materials wherewith men build hives for thee; then eat of every kind of
fruit, and walk in the beaten paths of thy Lord.’ There proceedeth from
their bellies a liquor of various colours, wherein is a medicine for
men. Verily herein is a sign unto people who consider.”
It is remarkable that the bee is the only creature that Mahomet assumes
the Almighty to have directly addressed. Al-Beidawi, the Arabic
commentator upon the Koran, whose authority ranks very high, in notes
upon passages of the preceding extract, says, “The houses alluded to are
the combs, whose beautiful workmanship and admirable contrivance no
geometrician can excel.” The “beaten paths of thy Lord,” he says, “are
the ways through which, by God’s power, the bitter flowers, passing the
bee’s stomach, become honey; or, the methods of making honey he has
taught her by instinct; or else the ready way home from the distant
places to which that insect flies.” The liquor proceeding from their
bellies, Al-Beidawi says, “is the honey, the colour of which is very
different, occasioned by the different plants on which the bees feed;
some being white, some yellow, some red, and some black.” He appends a
note to where Mahomet says, “therein is a medicine for man,” which
contains a curious anecdote. The note says, “The same being not only
good food, but a useful remedy in several distempers. There is a story
that a man once came to Mahomet, and told him his brother was afflicted
with a violent pain in his belly; upon which the Prophet bade him give
him some honey. The fellow took his advice; but soon after, coming
again, told him that the medicine had done his brother no manner of
service. Mahomet answered: ‘_Go and give him more honey_, for God speaks
truth, and thy brother’s belly lies.’ And the dose being repeated, the
man, by God’s mercy, was immediately cured.”
That the primitive Egyptians were familiar with the peculiar economy of
the bee in its monarchical institution is proved by the figure of the
bee being adopted as the symbolical character expressive of the idea of
a people governed by a sovereign. This figure is frequently met with
upon Egyptian sculptures and tablets, dating as far back as the twelfth
dynasty; but upon these the bee is very rudely represented, being
figured with only four legs and two wings; but upon a tablet of the
twentieth dynasty the bee is correctly represented with four wings and
six legs.
All these facts take us far back in the history of the bee. But the
indication of a higher antiquity of its domestication may be traced in
the Sanskrit, wherein _ma_ signifies honey, _madhupa_, honey-drinker,
and _madhukara_, honey-maker, the root of the latter signifying “to
build.” _Madhu_ has clearly the signification of our _mead_, thence we
may thus trace an affinity, pointing to those early times, for the
origin of a drink still in use amongst us. In Chinese _mih_, or _mat_
(in different dialects) signifies honey, thus clearly showing a second
derivation, in this Turonian term, from a more primitive language whence
both flowed. In the Shemitic branch nothing analogous is to be traced.
But this double convergence to a more distant point veiled in the
obscurity of time, necessarily takes the domestication of the bee back
also to that anterior period now only dimly traceable.
There can be but little doubt that the majority of the creatures now
domesticated by man were in those ancient days subjected to his sway,
and to which later times have not added any, or but few fresh ones. A
natural instinct possibly prompted him originally in the selection; and
if the reindeer of the Laplander seem an aberration, this has happened
through the contingency of climate, for in the high latitudes it
inhabits, it, in its uses to man, supplies the double function performed
in more southern regions by the equine and bovine tribes.
In the Greek and in the Teutonic languages, two branches of the Aryan
stem, the names of the bee, _melissa_ and _biene_, are clearly derived
from the constructive faculty of the insect, and to which the root of
the Sanskrit word _madhukara_, above noticed, also points. It would
seem, therefore, that an earlier notice of its skill than of its honey,
had suggested its name. Thus everything points to a very early
acquaintance with the bee, its economy, and its properties, and this
familiarity might be easily traced down in regular succession to the
present times, were it desirable to recapitulate what has been so often
repeated in the history of the “Honey-bee.” The facts I have gathered
together above, do not seem to have been hitherto strung together, and
may be suggestive of reflection, as well as affording some amusement.
-------
The study of the geographical distribution of natural objects has a more
universal bearing, and yields collectively more definite instruction and
information than its partial treatment, when restricted to small groups,
may at first seem to promise. This, however, is very useful, for it is
but by the combination of such special details that the enlarged views
are to be obtained, from which theories of the general laws of
distribution can be deduced. Of course, small creatures with locomotive
capacities will not supply the positive conclusions that may be framed
from such objects as are fixed to their abode, and have not the same
power of diffusion, although they certainly appear to be generally
restrained within particular limits by physical conditions of the
earth’s surface subservient to the maintenance of special forms of
organic life; and these, once determined, would yield and derive
reciprocal illustration. They may be merely climatic, but climate thus
indicated cannot be estimated by zones, or belts, or regions; for they
seem to traverse all these, and follow undulations not specially
appreciable except in the results they exhibit.
Unfortunately the bees have been too imperfectly collected, and too
irregularly registered, to admit of arriving at any precise conclusions
with respect to them. All that can as yet be done will be to combine the
scanty notices afforded by the contents of our collections, in the hope
that their promulgation may induce collectors, who happen to have the
often extremely rare opportunity of examining distant countries, to
avail themselves of the happy chance, which may never recur, or only at
long intervals.
Nor can I too impressively reiterate the importance of noting both
special localities, altitude, temperature, season, flora, etc., as being
all conducive to the widest instruction upon the subject. Indulging in
the hope that travellers will act upon these suggestions, and thus
considerably add to the value of what they may industriously collect, we
must patiently await until time brings it about.
Encouraging this expectation, I have summarily collected, under their
topical arrangement, the notices which precede, but which are there
arranged in the generic order of the bees.
From the information we thus possess, we learn that some of our genera
have an extremely wide diffusion, and occur in countries where we might
have expected that other forms would have superseded them in the offices
they are ordained to fulfil. None of the schemes for the geographical
distribution of insects yet propounded, seem to curb the eccentricities
of their range. The regions proposed by Fabricius in his ‘Philosophia
Entomologica,’ they break through as readily as through the concentric
circles of the cobweb when this opposes them: and all I can do is to
present them as they offer themselves, with the remark that the
occurrence of solitary forms in certain localities are almost sure
indications that allied genera would be found at hand were they
heedfully sought. It will also be observed, that in some places a
parasitical genus, and its known sitos, only, have been captured there.
The following list will strongly show how totally our genera of bees are
unaffected by isothermal, isotheral, or isocheimal lines drawn over the
earth’s surface. Nor do botanical conditions seem to influence them
beyond, the probability of their dissemination being restricted to the
special diffusion of the families of such plants whose genera and
species they frequent with us.
Thus, inhabiting Northern Europe we find in—
_Lapland._ Cilissa; Anthophora; Epeolus; Megachile; Chelostoma;
Heriades; Osmia; Apathus; Bombus; Apis.
_Finland._ Colletes; Prosopis; Cilissa; Anthophora; Nomada; Epeolus;
Stelis; Cœlioxys; Megachile; Anthidium; Chelostoma; Heriades;
Osmia; Apathus; Bombus; Apis.
_Sweden._ All our genera except Sphecodes; Halictus; Macropis;
Anthocopa.
_Denmark._ All our genera except Macropis and Anthocopa.
_Russia._ All our genera except Macropis and Anthocopa.
_The other Northern European Countries._ All our genera, with the same
exceptions.
Western, Southern, and Eastern Europe present us with, in—
_France._ All our genera.
_Portugal._ Prosopis; Sphecodes; Andrena; Halictus; Eucera; Nomada;
Anthidium; Apathus; Bombus; Apis.
_Spain._ Prosopis; Sphecodes; Andrena; Halictus; Dasypoda; Eucera;
Anthophora; Nomada; Megachile; Anthidium; Apathus; Bombus; Apis.
_Italy._ Andrena; Halictus; Panurgus; Eucera; Anthophora; Nomada;
Melecta; Epeolus; Cœlioxys; Megachile; Anthidium; Osmia; Apathus;
Bombus; Apis.
_Sicily._ Prosopis; Sphecodes; Eucera; Anthophora; Melecta; Epeolus;
Megachile; Anthidium; Osmia; Apathus; Bombus; Apis.
_Malta._ Halictus; Apis.
_Isles of Greece._ Dasypoda; Apis.
_The Morea._ Prosopis; Sphecodes; Halictus; Dasypoda; Eucera;
Anthophora; Ceratina; Nomada; Melecta; Anthidium; Chelostoma;
Osmia; Bombus; Apis.
_Albania._ Prosopis; Sphecodes; Dasypoda; Eucera; Ceratina; Nomada;
Melecta; Megachile; Anthidium; Osmia; Bombus; Apis.
_Dalmatia._ Halictus; Eucera; Anthophora; Megachile; Anthidium; Apis.
Asia exhibits to us, in—
_Siberia._ Andrena; Nomada; Epeolus; Bombus; Apis.
_Kamchatka._ Bombus.
_China._ Halictus; Nomada; Anthophora; Megachile; Bombus; Apis.
_Northern India._ Prosopis; Sphecodes; Andrena; Halictus; Ceratina;
Nomada; Cœlioxys; Megachile; Bombus; Apis.
_Bengal._ Anthophora; Ceratina; Apis.
_Tranquebar._ Nomada; Apis.
_Ceylon._ Anthophora; Ceratina; Apis.
_Bombay._ Anthophora; Megachile; Apis.
_Arabia Felix._ Anthophora; Anthidium; Apis. NOTE.—The genus _Apis_
does not occur in _Oman_.
_Mesopotamia._ Eucera; Nomada; Melecta; Megachile; Anthidium.
_Syria._ Halictus; Eucera; Anthophora; Cœlioxys; Anthidium; Bombus;
Apis.
In Africa we find, in—
_Egypt._ Colletes; Sphecodes; Andrena; Dasypoda; Eucera; Anthophora;
Saropoda; Cœlioxys; Anthidium; Osmia; Apis.
_Nubia._ Anthidium; Anthophora; Apis.
_Abyssinia._ Megachile; Apis.
_Tunis._ Dasypoda; Nomada; Apis.
_Algeria._ Colletes; Prosopis; Sphecodes; Andrena; Panurgus; Eucera;
Anthophora; Ceratina; Nomada; Cœlioxys; Megachile; Anthidium;
Osmia; Bombus; Apis.
_Barbary._ Halictus; Nomada; Anthidium; Osmia; Apis.
_Madeira._ Halictus; Apis.
_Canaries._ Colletes; Sphecodes; Andrena; Halictus; Anthophora;
Melecta; Osmia; Apis.
_Senegal._ Halictus; Anthophora; Ceratina; Megachile; Apis.
_Gambia._ Cœlioxys; Megachile; Anthidium; Apis.
_Sierra Leone._ Halictus; Cœlioxys; Megachile; Anthidium; Apis.
_Coast of Guinea._ Anthophora; Cœlioxys; Megachile; Anthidium; Apis.
_Fernando Po._ Megachile.
_Western Africa._ Halictus; Apis.
_Cape of Good Hope._ Halictus; Anthophora; Ceratina; Epeolus;
Cœlioxys; Megachile; Anthidium; Apis.
_South Africa_ [no distinct locality]. Halictus; Saropoda; Apis.
_Natal._ Anthophora; Cœlioxys; Megachile; Anthidium; Osmia; Apis.
_Madagascar._ Apis.
_Réunion._ Halictus; Apis.
_Mauritius._ Megachile; Apis.
In America we find, in—
_Arctic America and Hudson’s Bay._ Prosopis; Andrena; Halictus;
Megachile; Osmia; Bombus.
_Canada and Nova Scotia._ Andrena; Halictus; Nomada; Cœlioxys;
Megachile; Osmia; Bombus.
_United States._ Colletes; Sphecodes; Andrena; Cilissa; Halictus;
Eucera; Anthophora; Ceratina; Epeolus; Stelis; Cœlioxys;
Anthidium; Chelostoma; Heriades; Osmia; Apathus; Bombus.
_Mexico._ Anthophora; Epeolus; Megachile; Anthidium; Bombus.
_California._ Bombus.
_Columbia._ Colletes; Bombus.
_Quito._ Bombus.
_Chili._ Sphecodes; Halictus; Anthophora; Melecta; Epeolus; Anthidium;
Bombus.
_Jamaica._ Halictus.
_Cuba._ Anthophora; Cœlioxys; Megachile.
_St. Domingo._ Anthophora.
_Antigua._ Bombus.
_Guadeloupe._ Anthophora.
_St. Thomas’s._ Cœlioxys; Megachile.
_St. Croix._ Megachile.
_Cayenne._ Halictus; Eucera; Ceratina; Cœlioxys; Anthidium; Bombus.
_Pará._ Anthophora; Cœlioxys; Bombus.
_Brazils._ Prosopis; Halictus; Ceratina; Epeolus; Stelis; Cœlioxys;
Megachile; Anthidium; Apathus; Bombus.
_Paraguay._ Anthophora.
_Monte Video._ Bombus.
In Polynesia there occur—
_Sandwich Islands._ Prosopis.
_Philippines._ Anthophora; Nomada; Megachile.
In Australia are found—
_Swan River._ Prosopis; Megachile.
_Adelaide._ Prosopis; Megachile.
_Port Phillip._ Prosopis.
_Tasmania._ Prosopis; Megachile.
_Sydney._ Sphecodes; Halictus.
_New Zealand._ Halictus.
_Australia_ [but no distinct locality]. Anthophora; Saropoda.
------------------------------------------------------------------------
CHAPTER IV
NOTICE OF THE MORE CONSPICUOUS FOREIGN GENERA OF BEES.
SEEING thus the wide and almost universal distribution of many of our
own genera, we might be induced to ask whether this could not suffice,
by the impetus which more genial climates give to the multiplication of
individuals, to meet all the exigencies of the most favoured regions of
the vegetable kingdom. This is not so. There seems scarcely a limit to
the exuberance wherein nature revels in the production of variations of
form. The splendour, elegance, and infinite variety which she displays
in her floral beauties in the most luxuriant climates, find rivalry as
well in the multitude as in the magnificence of the insects which she
has allied with them as the indispensable promoters of their
perpetuation. How otherwise than through some of the insects we shall
mention could tropical _Labiatæ_ and the tubulated flowers of the
_Rubiaceæ_, etc. be fertilized? The reader will therefore, I trust,
welcome an acquaintance with some of the most conspicuous of the group
of bees produced by tropical countries, although the main object of this
treatise is to exhibit the attractions of “our native bees.”
I will but superficially and rapidly glance at the more distinguished
exotic genera and species, as supplementary to the preceding notice of
the geographical range of those which are indigenous with us.
How our own species reached us is a subject which has at present eluded
all satisfactory determination. For its solution we must await the
further discoveries of geology; at present we can only attribute their
advent here to the same causes which are common to the production of all
our groups of both the animal and the vegetable kingdoms.
Knowing how affluent tropical and sub-tropical countries are in the
variety, size, and number of the forms, as well as in the splendour of
their plants and vertebrated animals, we may fairly expect as gorgeous a
richness in the insects they produce. Nor shall we be disappointed, for
the imperial magnificence of their _Lepidoptera_ and _Coleoptera_
guarantees an equivalent brilliancy in the other orders of insects, and
which is fully confirmed by the harmonious splendour of their bees.
They thus put forward claims to attention and must excite curiosity by
their beauty and size, which the comparative smallness of our own, and
the usual dulness of their colours do not possess. The latter only repay
notice upon close investigation, but they then as amply reward all
labour bestowed upon them by the mental recreation they yield, as their
more gaudy exotic rivals. The former present themselves obtrusively and
exact notice, whereas ours meekly solicit it by their humble but solid
allurements. Here, as well as there, we behold the works of a mighty
hand and of an immeasurable intelligence.
The bees throughout the world, as known collectively to the richest
cabinets, number about two thousand species. This host, in itself
numerically so large, solicits attention, for it is opposed to the
economy of nature that there should exist any without functions of
essential usefulness, making them important elements in her harmonious
order and necessary to her due course, irrespective of the instruction
to be derived from the study of the manifold varieties of structure,
which unquestionably point to distinguishing peculiarities of habits.
In the true bees the division of the _Dasygasters_ presents the fewest
differing generic forms: the _Nudipedes_ and _Scopulipedes_ exhibit more
numerous varieties, the preponderance being in favour of the
pollen-collecting bees (the latter), although the cuckoo-bees (the
_Nudipedes_) are very abundant, and taken _en masse_, are certainly the
handsomest. If it be absolutely the case that there are no parasites
amongst the _Andrenidæ_, this subfamily will add very largely to the
exotic pollinigerous majority, which thereby becomes extensively
subservient to the fruition of the vegetable kingdom.
Those bees which are exclusively inter- or sub-tropical, seem furnished
with larger capacities for fulfilling the special mission to which the
family is appointed. Their pollinigerous and honey-collecting organs are
peculiarly adapted both to the structure and luxuriance of the superb
vegetation of those regions, and to which they seem distinctly limited.
But that they are not considered equivalent to the entire demand of the
profuse bloom everywhere abounding, may be concluded from the tropical
range and distribution of many of our northern forms. Thus, whilst the
flora of those climates is strictly circumscribed in its diffusion, its
fauna, distinctly in the class of insects, and especially in the family
of bees, is very considerably less limited, in extension.
The exotic genera of bees which are peculiarly noticeable, either from
splendour, size, or remarkable eccentricities of structure, are
numerous. Tropical and sub-tropical regions of course abound with them,
in individuals, in species, and in genera; and when we reflect upon the
riches of the flora of those countries, which is perpetuated mainly by
the agency of insects, amongst which, in fulfilling this indispensable
demand, bees, as I have reiterated, are pre-eminently conspicuous, we
shall not even wonder that their number, although excessive in the
extreme, is considerably aided, in many cases, in the performance of
this task, by peculiarities of structure. Thus, the splendid Brazilian
genus _Euglossa_, although not conspicuous for size, is remarkably so
for the enormous development of its posterior tibiæ, which form very
large triangles, compared with the size of the insect, deeply hollowed
for the conveyance of pollen. Its tongue also, from the length of which
the genus derives its name, is, when extended, more than twice the
length of the body, and with which it is enabled to reach the nectarium,
seated within the depths of the longest tubes of flowers. Other exotic
bees, further to aid them in collecting pollen, in addition to the dense
brushes with which their posterior legs are variously covered, have each
individual hair of these thick brushes considerably thickened by hairs
given off laterally, and in some cases these again ramify. Sometimes, in
variation, the simple, single hairs have a spiral curve, which almost
equally enlarges the activity of their operation. This is also the case
with two very hairy-legged genera of our native bees, proximately allied
to each other in the methodical arrangement, _Dasypoda_ and _Panurgus_,
the hair of whose posterior legs have this spiral twist. The most
hairy-legged exotic bees are essentially the genera _Centris_ and
_Xylocopa_. Of the habits of the former we know nothing, but those of
the latter we are intimately acquainted with, through the elaborate
descriptions given by Réaumur and the Rev. L. Guilding, the latter of
whom made his observations upon a species found in the island of St.
Vincent’s, in the West Indies. This last genus exhibits in some of its
species the giants among the bees, and one is especially so, a native of
India, the _Xylocopa latipes_, which is an inch and a quarter long, and
more than three inches in the expansion of its black, acute wings; and
it is also noticeable from the anterior tarsus in the male being greatly
dilated and white, the bee itself being intensely black, and which in
this same sex has enormous eyes united at the vertex, as in the male
_Apis_, or drone. In this genus, as in many other genera of bees, there
is often a great discrepancy in the appearance of the sexes, they being
so totally dissimilar that no scientific skill has hitherto been able to
discover a clue for uniting together correctly, by scientific process
merely, the sexes of a species; thence the numbers of the species in
such genera are unduly augmented beyond their natural limits, from the
fact of observation having neglected to associate the legitimate
partners.
In some of our native genera this same difficulty existed, which,
however, is gradually diminishing as the authentic sexes are slowly
discovered.
Exotic bees exhibit also a peculiarity I had occasion to observe before,
in reference to our own bees, amounting perhaps to a law, viz the more
highly-coloured condition of the parasite, for we find all the
parasitical bees of those latitudes, usually gorgeously arrayed in
metallic splendour, as instanced in _Aglaë_, _Mesonychia_, _Mesocheira_,
etc., and _Melissoda_ (my _Ischnocera_, in Lardner), is remarkably
conspicuous for its long and delicately slender antennæ in the male,
each joint of which is nodose at its extremity.
The widely-distributed _Nomia_ seems to abound chiefly in India. It,
although neither gay nor large, has, in its males, a distinguishing form
of the posterior tibiæ, which is greatly incrassated or thickened; a
peculiarity of structure found also in some other genera of
_Hymenoptera_, and in several genera of the _Diptera_, giving the
insects which have it a remarkable gait.
The singularly anomalous distortion of these posterior legs is
conspicuous also in the genus _Ancylosceles_, which is named in allusion
to it.
Another remarkable peculiarity is to be observed in the above genus,
_Mesocheira_, as likewise in the superb _Acanthopus_, both of which
genera have the spur of the intermediate leg palmated at the extremity,
and the latter genus is further distinguished by its large size and
splendid development, and by having the fifth joint of the tarsus of the
posterior legs longer than the three preceding united, and covered with
a pollinigerous brush as dense as that of the elongate first joint of
the same limb.
But the foreign genera which will be most interesting to the reader
will, I expect, be those of _Trigona_ and _Mellipona_, which, in many
peculiarities, seem abortive _Apes_. They seem nature’s first endeavour
to construct _Apis_, for they have an apparently imperfect neuration of
the wing, in which the external submarginal cell is unfinished. Their
only separating distinction from each other is the difference in their
mandibles, which in _Mellipona_ are broad and edentate, whereas in
_Trigona_ they are also broad but denticulated. In _Apis_ these organs
are merely irregularly enlarged at the extremity, and hollowed within,
rather like a spoon, which structure would of course imply a difference
of economy.
A further characteristic of these genera, and in which they participate
with _Apis_, is the deficiency of spurs to the posterior tibiæ, which
separates them from all other genera of bees, as also from _Bombus_,
which has two, yet with which, in point of their economy, they more
closely assimilate than with _Apis_. They are the South American and
Australian indigenous representatives of the genus _Apis_, and are found
likewise in Java and Sumatra, and in some of the larger and extreme
islands of the Indian Archipelago, thus also similarly in countries
where marsupial animals occur. Like _Apis_, they are social in their
habits; but their neuters only are as yet known, neither males nor
females having been described. They are reputed to be stingless, and to
make honey and wax in enormous quantities. The combs in _Mellipona_ are
attached either to the branches of trees or are suspended from them, but
how they are enveloped for security is not reported, but sometimes, like
_Apis_, they construct them within hollow trees and in the cavities of
rocks, as in _Trigona_, in like manner as _Apis_ does in its natural
state. Their communities are not so large as those of the hive bee, and
the cells of their combs are less perfectly hexagonal, the wax being
expended upon them in denser quantities, whereas the hive bee is
exceedingly parsimonious in the use of this material, a circumstance
arising possibly from the different and more difficult mode the latter
have of obtaining it. In the latter it is a secretion; but these exotic
genera possibly collect their wax ready-made by the exudation of plants,
and, thus, having more readily obtained it, they are more lavish in its
use.
Early travellers and historians describe many kinds of honey made by
these bees, native to the South American continent, but they report
nothing of the peculiarities of the social economy of these insects, nor
whether they are as closely allied in this respect to _Apis_, as they
are in the collection of honey and wax.
To enter into further detail relative to them would be beyond the
province of this work, and I have only given this extremely superficial
and brief notice of foreign genera, to show what multitudes of others of
this interesting family await admiration and study, when some
proficiency has been acquired in the knowledge of our own.
------------------------------------------------------------------------
CHAPTER V
PARASITES OF BEES AND THEIR ENEMIES.
NATURE seems to have imposed a restraint upon the undue increase of all
its creatures, by creating, to check it, others that prey upon them. It
thus enlarges the sphere of its activity by making life accessory to
life, and promoting thereby a more extended enjoyment of all its
pleasures. Other forms are brought into existence, and other terms given
to duration than those which the laws of life attach to specific
organization. No abatement is thereby made upon the quantity of
contemporaneous vitality, for what subsides in one rises in another, and
the undulation of the waves is perpetual.
Does the quantity of life, extant upon the earth, vary? Perhaps
mortality ever comes in some shape to prevent it, when excess threatens
to render its energy effete. Yet under every circumstance the wise
arrangements of Providence suffice, for everything has its enemies or
its parasites, which are also enemies, but frequently in disguise. For
defence there is an implanted instinctive fear, or abhorrence; and the
creature is then left to its skill, prudence, or strength, either to
evade or to mitigate, to the extent of its capability, the danger of the
attack.
We find the bees are not at all exempted from this prevailing condition.
They have many enemies and parasites of remarkably differing
organization. They are attacked by many kinds of birds, among which the
_Merops Apiaster_ (or bee-eater) is conspicuous. All the swallow tribe
prey upon them, as do the shrikes and some of the soft-billed small
birds, and also many small quadrupeds when they can find the
opportunity. Wasps also attack them, but they do not often get entangled
in spiders’ nets, being generally too strong for the retention of its
meshes, but I have seen a _Bombus_ enveloped in a tangle of its
wonderful filament.
The wild bees’ parasites are of two kinds, personal, and such which,
like the young of cuckoos, live at the expense of the offspring. The
personal parasites are again of two kinds, for bees are infested with
several kinds of _Acari_, and once I found a _Bombus_ upon the ground in
Coombe Wood so swarming with the _Acarus_ that it lay hopelessly
helpless until I threw it into a pool of water, when its _attachés_ were
washed away. But the poor bee seemed so prostrated by their attack, that
even when freed from them it had not energy to fly, and having landed it
I left it to the kindly nursing of nature.
A little yellow hexapod larva sometimes also infests the wild bees in
great numbers, running over and about them with great activity. I have
never followed these to their development, but they are said to be the
larvæ of _Meloe proscarabæus_, a conspicuously large coleopterous
insect. The assertion has produced much discussion; and I believe the
larva has been bred to the imago, and consequently it has been proved
that it is the larva of that insect. But that it should be parasitical
upon so small a creature, and that numbers should infest it for their
nutriment, is extremely improbable. It is far more likely that instinct
has taught them to be conveyed elsewhere through the medium of the bee,
as they might also be by attaching themselves to any other volatile
insect, and that upon arriving at a suitable locality they descend from
their temporary hippogriff. We see seeds thus conveyed by the agency of
animals and birds to suitable places, where they fall and germinate.
Another little hexapod is occasionally found upon them: this is
intensely black, and like the former, very active: these I never could
rear, nor did they ever seem to enlarge, and they speedily died. I have
found them in profusion also within the flowers of _syngenesious_ or
composite plants, especially of the dandelion in the spring.
But their most remarkable personal parasites consist of some very
extraordinary insects, so anomalous in their structure as to have
required the construction of an order for their reception,—the Order
_Strepsiptera_, or “twisted-winged,” thus named from the twist taken by
their anterior wings or wing-cases. Their natural history is but
imperfectly known, and I believe the males have not yet been discovered.
Their larva lives within the bee, and feeds on its viscera by
absorption, being attached within by a sort of umbilical cord. It
presently consumes the viscera, and renders the bee abortive, by
destroying its ovaries, for it is usually upon female bees that it is
found. When full fed it forms a case within which it changes into the
pupa and imago, the head of which case protrudes between the scales of
one of the dorsal segments of the abdomen. How it becomes deposited
within the bee or the bee’s larva remains a mystery, although many
hypotheses have been hazarded to account for it, but all are
unsatisfactory. The Order consists of three genera (_Stylops_,
_Elenchus_, and _Halictophagus_) found in England, and other parts of
Europe; indeed, the genus _Elenchus_ has been also discovered in the
Mauritius. The Continent possesses the genus _Xenos_, of the same order,
and parasitical upon a wasp, neither of which occur with us.
Mr. Kirby, in studying the bees for his invaluable ‘Monographia Apum
Angliæ,’ first came across this extraordinary creature. His description
of his discovery is highly interesting. He says, at page 111 of volume
ii. of the above work, that having observed a protuberance upon the body
of the bee, he was anxious to ascertain whether it might be an _Acarus_,
and goes on: “What was my astonishment when, upon attempting to
disengage it with a pin, I drew forth from the body of the bee, a white
fleshy larva, a quarter of an inch long, the head of which I had
mistaken for an _Acarus_. How this animal receives its nutriment seems a
mystery. Upon examining the head under a strong magnifier, I could not
discover any mouth or proboscis with which it might perforate the
corneous covering of the abdomen, and so support itself by suction; on
the under side of the head, at its junction with the body there was a
concavity, but I could observe nothing in this but a uniform unbroken
surface. As the body of the animal is inserted in the body of the bee,
does that part receive its nutriment from it by absorption? After I had
examined one specimen, I attempted to extract a second, and the reader
may imagine how greatly my astonishment was increased, when, after I had
drawn it out but a little way, I saw its skin burst, and a head as black
as ink, with large staring eyes, and antennæ consisting of two branches,
break forth, and move itself briskly from side to side. It looked like a
little imp of darkness just emerging from the infernal regions. I was
impatient to become better acquainted with so singular a creature. When
it was completely disengaged, and I had secured it from making its
escape, I set myself to examine it as carefully as possible; and I
found, after a careful inquiry, that I had not only got a nondescript,
but also an insect of a new genus whose very class seemed dubious.”
As everything connected with so strange a creature is very attractive, I
will cite what other observers also have seen. Mr. Dale, from whom
Curtis received _Elenchus_ to figure in his ‘British Entomology,’ vol.
v. pl. 226, says: “These parasites look milk-white on the wing, with a
jet-black body, and are totally unlike anything else. It flew with an
undulating or vacillating motion amongst the young shoots of a quickset
hedge, and I could not catch it until it settled upon one, when it ran
up and down, its wings in motion, and making a considerable buzz or hum,
as loud as a _Sesia_; it twisted about its rather long tail, and turned
it up like a _Staphylinus_. I put it under a glass and placed it in the
sun; it became quite furious in its confinement, and never ceased
running about for two hours. The elytra or processes were kept in quick
vibration, as well as the wings; it buzzed against the sides of the
glass with its head touching it, and tumbling about on its back. By
putting two bees (_Andrena labialis_) under a glass in the sun, two
_Stylops_ were produced: the bees seemed uneasy, and went up towards
them, but evidently with caution, as if to fight; and moving their
antennæ towards them, retreated. I once thought the bee attempted to
seize it; but the oddest thing was to see the _Stylops_ get on the body
of the bee and ride about, the latter using every effort to throw his
rider.
“As the _Stylops_ emerges from the body of the bee, the latter seems to
suffer from much irritating excitement.”
Mr. Thwaites writes to me, on the 12th May, thus: “I had the good
fortune to capture a _Stylops_ flying, and on the Tuesday following saw
at least twenty flying about in the garden, but so high from the ground
that I could capture only about half-a-dozen; since that time they have
become gradually more scarce.
“The little animals are exceedingly graceful in their flight, taking
long sweeps as if carried along by a gentle breeze, and occasionally
hovering at a few inches distance from the ground. Their expanse of wing
and mode of flight give them a very different appearance to any other
insect on the wing. When captured they are exceedingly active, running
up and down the sides of the bottle in which they are confined, moving
their wings and antennæ very rapidly. Their term of life seems to be
very short, none of those I have captured living beyond five hours, and
one I extracted from a bee in the afternoon was dead the next morning.
“All the bees stylopized, both male and female, I have taken, have
manifested it by having underneath the fourth (invariably) upper segment
of the abdomen a protuberance which is scale-like when the _Stylops_ is
in the larva state; but which is much larger and more rounded when the
_Stylops_ is ready to emerge. A bee gives nourishment generally to but
one _Stylops_; but I have occasionally found two, and once three larvæ
in one bee.”
The structure of these insects is very remarkable: the typical genus
_Stylops_ is named from its compound eyes, which consist of a very few
(about fifteen) hexagonal facets, seated upon a sort of footstalk. The
mandibles are lancet-shaped and very acute, and the head, by reason of
the protuberant eyes, has very much the shape of a dumb-bell. The
antennæ are branched, but in _Halictophagus_, they are flabellate. The
thorax is greatly developed; the superior wing is like a rudimentary
wing-case, and is twisted, the inferior wings are very large, and fold
along the abdomen in repose like a fan; the legs are slender, and the
tarsi with four joints in _Stylops_, with three in _Halictophagus_, and
with two in _Elenchus_; the abdomen is long, very flexible, and consists
of eight segments. The insects themselves do not exceed a quarter of an
inch in length in the largest, but they are generally very much smaller.
The perfect insect is very short-lived, not surviving many hours, as
just stated. They are usually found in the months of May and June, and
they have been discovered to infest several species of _Andrena_ and
_Halictus_, for instance the _A. nigro-ænea_, upon which Mr. Kirby first
found it; _A. labialis_, which I have frequently caught stylopized; _A.
rufitarsis_, _fulvicrus_, _Mouffetella_, _tibialis_, _Collinsonana_,
_varians_, _picicornis_, _nana_, _parvula_, _xanthura_, _convexiuscula_,
_Afzeliella_, _Gwynana_, etc., and upon _Halictus æratus_, etc.
The other mode of parasitism destructive to the bees is where the
parasite deposits its own egg upon the provender stored by the bee for
the sustenance of its own young. The young of the parasite, either by
being more speedily hatched or more rapacious than the larva of the
sitos, starves the latter by consuming its food. This kind of parasites
consists of several _Diptera_, but they are mostly bees which form a
distinctive subsection of the family of true bees (_Apidæ_), the
subsection being called the _Nudipedes_ or naked-legged, from their not
having the necessary apparatus of hair upon the posterior thighs or
shanks, for the conveyance of pollen wherewith to store their nests.
Thus nature, having rendered them unable to perform this duty to their
offspring, has imposed upon them the necessity of resorting to strangers
to support them, and they are not led to it by idleness or indifference.
These insects consist, with us, of six genera, the species of which are
individually attached to some particular bee, who thus nurtures their
young. They are, as a rule, gayer insects than those which they infest,
and the genus most abundant in species is _Nomada_, which attaches
itself chiefly to _Andrena_, although some of its species, especially
the smaller ones, infest the species of _Halictus_, and one frequents
_Eucera_. _Melecta_ appears confined to _Anthophora_; _Epeolus_ to
_Colletes_; _Stelis_ perhaps to _Osmia_, judging from the great
similarity of habit; and _Cœlioxys_ to the constructive _Megachile_.
None of these parasites resemble their sitos, but _Nomada_ is
exceedingly different, being in its gay array more like a wasp than a
bee. The only close approach in the appearance of a parasite to the
insect upon which it is parasitical is in the resemblance between
_Apathus_ and _Bombus_, which are so alike that they were long continued
to be united in the same genus, until the peculiar characteristic of the
parasitical bees was detected, when they were readily separated.
Although, cuckoo-bees as they are familiarly called, they could not be
associated with the _Nudipedes_, because their posterior legs, though
not pollen-conveying organs, are hairy; but the _Cenobites_, to which
section they belong, have a peculiar and distinguishing structure of
that limb. They are further separated from the _Nudipedes_ by several
frequenting the same nest, thus habitually associating with their sitos.
Some of the _Chrysididæ_ are likewise, as I shall have occasion to
notice in the description of the habits of the genera, similarly
parasitical upon some of the species of the family of bees. The genus
_Mutilla_ is also probably entirely parasitical upon bees, for _Mutilla
Europæa_ is a parasite upon _Bombus lapidarius_, from whose nests it has
been dug in winter, by my friend the late Mr. Pickering, whose activity
and accurate observation once promised to be very beneficial to the
science, but he, like many others of my entomological friends, is now no
more!
------------------------------------------------------------------------
CHAPTER VI
GENERAL PRINCIPLES OF SCIENTIFIC ARRANGEMENT.
THE following rapid observations are addressed to those whom it is the
desire that this series of volumes may induce to take up the study of
Nature in a methodical manner. With this view, the merest summary of the
principles upon which scientific arrangement is based, is here
exhibited. The study requires method as a lodestar to guide through its
intricacies, but it is one which, pursued simply as a recreation, yields
both much amusement and gratifying instruction. It shows us that when we
unclasp the book of nature, and wherever we may turn its leaves, every
word, the syllables of which we strive to spell, is pregnant with the
fruitfulness of wonderful wisdom, whose profound expression the human
intellect is too limited thoroughly to comprehend.
Is there an arrangement that human skill could mend? Is there an
organization that man can fully solve, or a combination that his mind
can wholly compass? Do we not behold limitless perfection everywhere,
but all so deeply mysterious. So exquisite are the feelings which the
contemplation commands, that they imbue us deeply with the sense of the
high privilege conferred upon the intellect by its being permitted to
embrace a study, which, even pursued merely as a relaxation, inculcates
in so serene and pleasing a manner such profound veneration and
reverence.
To acquire the prospect of a possibility to unravel the exuberant
profusion of the natural objects surrounding us, successive students of
nature have endeavoured to systematize the seeming confusion in which
her riches are spread about. Like has been brought to like, and
gradation made to succeed gradation. Resemblances have been combined and
disparities disjoined, until the labour of centuries has constructed of
all the natural objects within the ken of man a vast and towering
edifice, whose basis is seated at the lowest substructure of the earth
which research has yet reached, but whose head ascends high into the
empyrean.
All things have been collected, and arranged, and classed. Method has
endeavoured to give them succession according to an assumed
subordination. The labour of the great minds which framed the large
theories of this vast branch of human knowledge, has permitted men of
lesser powers of combination to abstract parts for special examination
and investigation.
The study of natural science has progressively reached an extraordinary
development, spreading in every direction its innumerable tentacula; to
which the perfection of the telescope and of the microscope have still
further added by the discovery of new worlds of wonder.
Just as language is systematized and made easier by grammar methodizing
its co-ordinates and their relations, so natural science arranges its
subjects into subdivisions of which genera and species are the lowest
terms. The higher and more complicated are of many denominations, which,
notwithstanding, have for their chief purpose the simplification of the
survey by assisting accurately to determine accurately natural objects
individually. Once the clue of the labyrinth caught, the seeming
intricacy of its involution vanishes; for when a clear conception of the
general scheme is obtained, the solution of the parts is comparatively
easy. The same principle rules throughout, however variously treated.
The large divisions of nature appear simple and distinct enough in their
great frame, but when we approach their confines, close investigation
discovers analogies and affinities, which, where the separation seems
most apparent, create insuperable difficulties, and render linear
succession, or distinct division, nearly an impossibility. _Here_ we
find parallelism, and _there_ radiation, and elsewhere a complicated
reticulation without subordination; and this is one of the great
problems, which it is the office of the mature naturalist to endeavour
to solve. The present work has to do, however, with but one small
portion of the whole.
Thus we see that, in order to arrive at a knowledge of natural objects,
a method must be pursued to avoid being overwhelmed by their
multiplicity, whereby confusion would be produced in the mind which
their methodical investigation tends to dissipate. Their abundance
precludes the possibility of their being all equally well known,
although it is very desirable to have a general, if even superficial
acquaintance with them, that is to say, in the broad and distinguishing
features of their large groups, for as to an accurate knowledge of all
their species, it would be futile to attempt it. Possessing this general
knowledge, the attention may be turned with greater advantage in any
special direction, and that pursued to its entire acquisition.
Natural objects have been arranged in KINGDOMS, ORDERS, CLASSES,
FAMILIES, and GENERA, all deduced in their successive and collateral
groups from characters exclusively derived from SPECIES; therefore to
the accurate knowledge of _species_ all endeavours must be directed,
they comprising within themselves all the rest, although the characters
upon which they themselves depend for separation from their congeners
are the most trivial of any. Each combination, in its analytical
descent, contains characters of wider compass than those which succeed
it, and consequently embraces in that descent more species than the
successive divisions; just as in the ascent, or synthetical method, the
characters of every successive group gradually expand. Species being
thus the only real objects in nature from which all knowledge springs,
and in which exclusively all uses lie, other combinations being perhaps
as merely imaginary as are the many lines which are drawn over the
surface of the globes, it would imply that subdivisions merely lend aid
to acquire more rapidly the details upon which they depend. We will,
therefore, first turn our attention to species.
Both combination and subdivision are intended to facilitate
identification, by aiding us to arrive at this knowledge of species; for
each species represents a distinct idea, whose correct definition is
important to the progress of accurate science. This alone permits
observation to be attributed to its right object, and when properly
recorded, the information is secured for ever from error or obscurity.
It is not, however, the gift of every mind to discern accurately even
specific differences, or to form skilfully generic combinations. The
very best favoured by nature,—for it is a natural gift, although under
high cultivation,—have sometimes a bias towards seeing more than
actually exists. Hence varieties are often elevated into species, and
species thus overwhelmingly multiplied; and genera are frequently framed
upon vague distinctions.
Species are the basis of all natural science.
A species in zoology is a combination of creatures which unites the
sexes, and these being two, the assumed existence of neuters in some
instances does not invalidate this, it comprises two individuals having
independent existence, but whose co-existence is indispensable to
perpetuation, but which often, from their great differences, no single
set of scientific characters will bind together, yet which must exist in
some undiscovered peculiarity, that individuals may be able to
distinguish their legitimate partners. The species, therefore, is a
complete unit in its entirety, although consisting of two distinct
beings, for in the large majority of cases in zoology these sexes are
distinct, although their conjunction is, in the higher forms of life,
indispensable for their continuance. In some of the lower forms of
animal life they exist in union, and in the vegetable kingdom we
perceive every possible combination and modification of this
conjunction, and in both of these life may be perpetuated also by
simpler processes.
The species may consist of any indefinite number of individuals, and no
law has hitherto been discovered which regulates the relative
proportions of the sexes, although it is very apparent that some
recondite influence operates to control it. It is also extremely
remarkable to observe how eccentric nature is in some species, and the
extent to which she sometimes carries the variation of some particular
specific type, and to which some species are singularly prone, and yet
how rigidly in other cases she adheres to the particular specific form
in the succession of generations, that even the shadow of a deviation
from the typical distinction is scarcely to be discovered: a reason for
this it is hard to surmise. We may, nevertheless, conclude it to be
certain that true species are ever distinct, and can no more coalesce,
however closely they may approach together, than can asymptotes.
Specific differences result from many characteristics,—from colour,
clothing, size, and sometimes from peculiarities of structure; but these
last are usually of a higher order, tending to indicate an aberration,
slight though it be, from the normal generic character which holds the
group together, thus implying a distinctive economy. This is sometimes
called a subgeneric attribute, and there might be a reason, certainly,
for not elevating such species to the full rank of genera, were genera
equivalents, which they are not, and it merely remains an evasive
admission of the doubt that attaches, except for the sake of
convenience, to any subdivision, but the specific.
The species is thus the very last term of subdivision, the very
elemental principle itself, which unites together as one, solely for the
purposes of perpetuation, the two sexes of similar individuals, and
without whose intercourse the kind or species would die out.
That some species greatly abound in individuals, as before observed,
whilst others appear to be extremely limited, is an absolute fact, and
not merely suggested by a defective observation of their occurrence,
resulting from their rapid dispersion. It is verified by being noticed
to occur where we know they would resort, as is exemplified in the case
of some of the parasitical species of the insects herein treated of, and
which are sometimes rare, even in the vicinity of the metropolis of
their sitos, and where this also greatly abounds. In other cases, other
species absolutely swarm where the similar attraction lies.
Even supposing species to be the sole natural division, we may accept
the superior combinations as means to aid us to a gradually extending
survey of the whole. Perhaps did we possess all the links of the vast
chain of beings we should find genera, and every other superior
combination, melt away through the intimate alliance of the succession
of species that would obliterate the lines of separation, by making the
sutures imperceptible; but what mind could compass the detail of such a
limitless unbroken series? Their subdivision may therefore be accepted
as a positive necessity, to enable us to compass their investigation. As
it at present stands, with our imperfect knowledge of the entire series
of species, these higher groups are indispensably requisite.
The specific diagnosis being the only sure basis upon which all our
knowledge can rest, its accuracy is all-important, and requires a few
observations. It comprises two parts—the specific _character_, and the
specific _description_. The difference between these is, that the first
is constructed with the extremest brevity consistent with its utility,
is fluctuating and not permanent. The latter permits all the diffuseness
needful to embrace a full description of the creature.
The object of the first is to establish the _present_ identity of the
species amongst all its known congeners—those associated in the same
genus;—and that of the second to secure it in its perpetual identity,
and segregate it from all future and contingent discoveries. The
specific character admits, consequently, modifications to suit any
extension of the genus, and in fact exacts it at the hands of all who
describe new species. This many naturalists undertake without any
apparent consciousness of the scientific responsibilities that attach to
it, and whence results the confusion so much to be deplored, of the
synonymy that prevails, constituting, as it does, such a Dædalian
labyrinth. The describer of a new species is bound to cast around, and
endeavour to know all that has been previously done upon the subject of
the genus. He has to revise all the specific characters within the
genus, and mould them to those he introduces, and he must insert these
closest to their evident affinities. Thus, therefore, the describer’s
labour is not light, if to be of any value. The specific character,
although thus varying, becomes a permanent utility, and only so fulfils
its object,—that of rapidly showing, at a glance, the known species of a
genus, and thereby permitting the speedy determination of the identity
or distinctness of a compared object. If doubt should exist from this
brevity, the specific description is at hand to solve it, by the
amplitude and completeness of its details. Of course this mode of
treatment is only suitable to monographs, or portions of the science
discussed separately, and not to a general or universal survey.
The amount of toil thus saved to the describing naturalist, and to those
who wish to name their specimen, the experienced only can estimate. This
brevity of specific character is one of Linnæus’s terse and valuable
axioms, who limits its length to twelve words. The best examples, I
think, that I can adduce in entomology, of valuable and exemplary
specific descriptions, is Gyllenhal’s ‘Insecta Suecica’ which contains
exclusively a description of Swedish Coleoptera; Gravenhorst’s large
monograph of European Ichneumons; Erichson’s elaborate work upon the
Staphylinidæ; and our own Kirby’s ‘Monographia Apum Angliæ.’ Their
perfection consists in fulfilling thoroughly all the above conditions,
for if any doubt exist upon comparing your insect with their
descriptions, you may be fully assured yours is not identical. The only
drawback to the utility of Mr. Kirby’s book is that he had to deal with
insects variable in condition from many causes, and the variable state
of the insect that may have to be compared; his description has
evidently been made sometimes from a worn specimen, one that had been
exposed to wind and weather, and sometimes from an insect in fine
condition. Thus it is important that compared insects should be in an
identical state to substantiate the comparison,—a difficulty which this
family has specially to contend with, as these insects are more liable
than almost any others to vary, owing to their specific character
depending much upon pubescence, which is extremely subjected to many
modifying influences, for the tinges and positive colour of the hair
will much vary by exposure, as it is not possible always to capture a
bright individual.
Taking specific description thus practically in its full and wide sense,
it is requisite, for the purpose of avoiding repetition, that all the
characters of the superior combinations should be eliminated, leaving it
with those only which have not been thus absorbed, which now constitute
its sole remaining distinctive specific peculiarities. Every species
necessarily contains within itself, every character of every combination
in direct line above it, although these have been gradually abstracted
to form those several combinations which are arrived at successively in
the synthetical ascent. Analytically, species are the last but combining
element of all, although their most remote members. The whole system is
an ingenious contrivance for breaking down a complex multiplicity of
characters, to simplify the means of reaching all the collateral or
adjacent species, that we may be able to determine identity or
difference.
Entomology, and indeed natural history generally, uses three words, very
much alike, but very different in signification and application. These
are, _habit_, _habits_, and _habitat_. The habit is that peculiar
character of identity, that _je ne sais quoi_, which marks all the
species of a genus collectively, and which, in some cases, only the
trained eye can detect. It is then seen instantaneously, and forcibly
illustrates the extreme precision the study of the natural sciences
tends to cultivate. Their utility, also, as a discipline to the mind,
conjunctively with the keen accuracy which practice gives the sight, are
qualifications not lightly to be esteemed.
It is from such absolute control of detail that the most efficient power
of generalizing emanates, which, when it has once become habitual,
gives, from its rapidity, an almost instinctive facility, as its
inevitable concomitant, for both synthetical and analytical survey. The
mind thus becomes strengthened by vigorous exercise, and has always, for
every purpose, a powerful instrument at command, often used
unconsciously, but always effectively. Thus is _habit_, once correctly
perceived, ever retained.
The _habits_ are the peculiar manners and economy of a species; and the
_habitat_ is the kind of locality the creatures affect, such as hill or
plain, wood or meadow, forest or fell, hedgebank or decaying timber,
sand or chalk or clay, and ground vertical or horizontal; and the
_metropolis_ of a species—another term in use—is the centralization of
the general habitat where the insect either nidificates collectively
with its fellows, or, where, from any other cause, it may be found in
its season, usually in profusion. But good fortune does not always
attend the discovery of this locality.
It is by the acquired skill of perceiving habit, that a large and
confused collection may be sorted rapidly, or fresh captures immediately
placed with their congeners, without the necessity of going tediously
through all the descriptive characteristics. Incidental errors are
afterwards speedily corrected. It is then that the specific character
exhibits its utility by enabling us at once to distinguish the new from
the old.
The concentration and summary of the specific character is the name of
the species, or trivial name as it is sometimes called, which is, as it
were, the baptismal designation that attaches to it always afterwards,
and is contemporaneous with the introduction of the creature into the
series of recognized beings.
Upon the revival of the study of natural history, when learning dawned
after the night of the Middle Ages, much difficulty attached to the
imposition of discriminative names. The works of the ancients were
ransacked, and endeavours made to verify and apply the names they had
used. Ray published a vocabulary of such names. But the ancients never
studied natural history in the systematic way pursued by the moderns;
they did not want the skill, but they wanted the facilities. Anatomy and
physiology had not made the progress necessary to aid them in the
pursuit, and the assistance all these sciences obtain from optical
instruments was barred from them. The names they gave to natural objects
were vernacular names, which, like our own vernacular names, applied
rather to groups than to species, and have in consequence ultimately
become the names of genera. But this was the work of time, with which
discovery progressed. As these discoveries were made by the new
cultivators of natural history, they added them to those which they
resembled, by some brief distinctive character adapted to the momentary
exigency, such as _major_, or _minor_, etc.; and these additions were
constantly treated as varieties of the species, whose name headed the
list by the designation first adopted. Discoveries still continued,
which were compulsively arranged with the predecessors they most nearly
resembled, until resemblances vanished, and the boundaries fixed by the
assumed correct application of the names thus derived from the ancients
were passed, and there was an overflow on all sides.
To meet this difficulty, the new discriminative name had to be moulded
into a phrase to correct its exceptive peculiarities, and specific names
became descriptive phrases, the bulk of which no memory could retain,
and which usually were neither clear nor expressive. Thus genera were
continually treated as species, and species as numbered varieties, with
long distinguishing descriptive phrases.
So it remained till day dawned, and the great luminary of systematic
natural history rose with a bound to irradiate the obscurity of science
with his subtile and vivifying beams.
This was LINNÆUS, to whom we owe the binomial system, wherein, by means
of two words only (the generic or surname, and the specific or baptismal
name), the recognition of a species is perpetuated; for Linnæus truly
says, “_Nomina si nescis, perit et cognitio rerum_.”
By a law tacitly admitted, but universally recognized, for the sake of
securing to a name its intangibility, no two genera in the same kingdom
of nature may be named alike. There is, therefore, if this rule be
observed, no fear of similar names coming into collision in the same
province, and thus producing confusion. A ready means to prevent the
possibility of such mischance is the admirable work which has been
published by Agassiz, with the assistance of very able coadjutors, in
the ‘Nomenclator Zoologicus,’ which is a list of all the generic names
extant in zoology, exhibiting what names are already in use either
appropriately or synonymously in this great branch of the natural world,
and if this work receive periodically its necessary supplements and
additions, no excuse will remain for the repetition of a name already
applied. The most defective character in this laborious work, is the
frequent incorrectness of its etymology of the names of genera. It would
be, perhaps, without such aid, too great a labour to require of the
describing naturalist, or it might not be otherwise even practicable for
him, to ascertain whether the generic name he purposes to impose be, or
not, anticipated. The penalty of its being superseded is understood to
attach to the imposition of such a name, for the alteration may be made
with impunity, and thereby it becomes degraded to the rank of a mere
synonym.
Nomenclature has thus, by the happy invention of Linnæus, been made a
matter of the greatest simplicity, conciseness, and lucidity, and to
him, therefore, our gratitude is due.
An indispensable branch of nomenclature is _Synonymy_, which, briefly,
is the chronological list of the several names under which species or
genera may have been known. This diversity of names has originated in
several ways,—from indolence, or ignorance, or excessive refinement. The
views of systematists will differ in the collocation of creatures;
hence, sometimes what had been previously divided will be recombined, or
divisions into further groups be made of what had been before united.
Both processes will necessarily produce synonyms; the recombination of
what had been separated reduces the names of such groups to the rank of
synonyms of the old one from which they have been disjoined. In the
latter case the old name will be retained to the typical species merely,
and be also made a partial synonym of the names of the new generic
groups: or, indeed, it may happen that the same creature has been
described generically, unknowingly, by two different persons, about the
same time. By another recognized rule in nomenclature, the ‘law of
priority,’ the name given by the first describer is accepted, and the
other consequently falls to the condition of a synonym.
With respect to specific synonymy, many causes conduce to it; namely, an
imperfect description which cannot be clearly recognized, reducing it to
that category, with a mark of interrogation appended; subsequent
description when want of tact has not discerned the identity of the old
one; indolence in looking about for works upon the same subject;
inability to obtain access to books wherein they may be described, owing
either to their costliness or to their obscurity, or by lying buried in
some collapsed journal, or the poverty of our public libraries, etc.
etc. But however thus lost sight of, or wilfully ignored, the name still
retains vital elasticity, for the describer has not thereby lost his
rights, but revives to them with all due justice upon the cessation of
this coma. The really culpable among such describers are those who
neglect to look around them to ascertain what has been done, and this
course is sometimes illicitly adopted to obtain a fleeting and
meretricious fame, by the description of ostensibly new species, which
critical investigators soon detect to have been long since known and
very ably described.
Thus, a complete synonymy, which can almost only come within the
province of a monograph, would give, chronologically, the entire history
of a species under all the names it has been known by in the several
works in which it has been published. Nature is so uniform and stable
that Aristotle’s descriptions can be clearly recognized, therefore there
is no fear that whatever may have been synonymously, but yet correctly
recorded of the economy of a species, can possibly be lost when once
registered in the archives of science.
The working out of a correct synonymy is an ungrateful task of much
labour, for few appreciate it, and not many use it, although when
thoroughly elaborated it is so extremely valuable.
A further rule in nomenclature is, that the generic name must always be
a substantive; and it is always desirable that the specific name should
be an adjective. In the event of the imposition of a proper name, which
is sometimes done to record a private friendship, but improperly so, for
it is a distinction due only to promoters of the science, the genitive
form must be adopted.
The next grade in ascent from the species is invariably the GENUS, for
subgenera, like varieties in species, are not uniformly present, but are
mere contingencies, even if they do properly exist.
Why some genera abound in species and others are so limited is as
difficult to determine as the differing numerical abundance of
individuals in species. That long genera (genera numerous in species)
may be the result of natural selection, as Mr. Darwin surmises, and the
offspring of a common parentage, is contradicted, not merely by peculiar
although sometimes slight dissimilarities of habit, combined with size
and colour, but also if any lines of demarcation are to be admitted, it
is possible, were their generic similitude to be subjected to severe
test, they might present characteristics normally discrepant and
suggestive of further division, although the habit may be very like.
The generic grouping is effected by structural peculiarities, which are
essentially of a higher class than the characters of specific
separation, these being determined by colour, pubescence, sculpture,
etc. etc.; specific characters combining only individuals with such
peculiar inferior resemblances. The generic characters thus establish
groups of species allied only by such more general character and
similarity, but conjunctively of one permanent habit, although the
members of the genus may differ somewhat in habits, and so on of the
higher groups into which insects are collected, each group in its ascent
upwards presenting characteristics of a wider range than those of the
descending series. And so, by degrees, we rise until we reach the
characters which combine the whole order. The process is necessarily and
imperatively synthetical, for the whole foundation is based upon
species, and thence emanates the supposition that only species exist.
The _type_ of a genus is that species upon the characters of which the
genus was originally framed and named, and theoretically, however
generic groups may be subsequently divided to suit views or to meet
systems, the primitive generic type is assumed to retain the primitive
generic name. It is much to be doubted whether, in every case, the type
is the true pattern, or leader, or centre of the group called the genus;
nor is it likely if genera be natural groups. It has usually been
accident which has dropped upon the favoured species, and not a
well-calculated and thoroughly digested selection, and which, although
accepted, will require emendation or change if the whole collective
series should ever be obtained.
It is the necessary result of the imperfection of our intellect, and one
of the dominant conditions of overruling time, that one thing must
follow the other. It is, therefore, neither an expressed nor even an
implied inferiority that puts one species before the other in a generic
group; or one genus before the other in their successive order.
Affinities may lead both species and genera in varying directions,
although treated descriptively as of linear succession, in which order
they are usually arranged, but this is unavoidable and therefore not
derogatory. It is for the mind to conceive their radiation from a type,
or their parallelism with other forms, even in the connection of
affinity, and not merely of analogy, for the latter can be expressed
even in arrangement.
Thus encouragement attends the beginner at the very outset of his study,
and the prospect of a wide field for discoveries, in all directions,
lies open to him.
The FAMILY, after the GENUS, is the next natural group at which we
arrive, proceeding synthetically. Its characters, succeeding to those of
the ORDER, group together collectively the largest numbers of forms that
in their several combinations are the most nearly equivalents, and may
be almost paralleled in that quality to the alliance of species.
Ascending from species, the naturalist scarcely hopes to find in the
groups formed above them strict parallelism, although, to be logical, it
should be so, and, where the combinations are most natural, it is most
nearly so. Thus we do not again distinctly reach equivalents until we
arrive at these families, which from linking together associations
usually combined by an identity of instinct and functions, attach to
themselves greater interest, and form alliances pointed out by the
finger of nature itself, which are therefore exempted from the arbitrary
caprice of the constructive systematist.
It does not follow that families should be even nearly numerically
equivalent, for a family may contain a few or a multitude of genera and
species, or a multitude of genera and few species, or also a multitude
of species and few genera. Families comprise groups of forms to which
nature delegates the execution of certain duties and offices, and
whether specifically numerous or few, we may assume they are sufficient
for the object intended. If we can reach the motive that controls the
peculiarities of the group, it is a golden key to the explanation of the
structure of its constituents, and, perhaps might furnish us, if not
with a positive clue, yet with a surmise as to the functions of the
collateral groups of which it forms a member, and which diligent
observation may accurately determine.
Families, to be natural divisions, should stand in the same relationship
to genera as species do, but from the opposite side, whatever the
subdivisions are into which they may be separated, for the sake of
convenience, and as descending grades whereby to arrive with greater
facility at their genera, just as the species of the latter are also
sometimes grouped, that they may be reached with greater ease. These
subdivisions of families have no analogy with the varieties which
species occasionally throw off, although they may be as irregular in
their occurrence; that is to say, in the association of a group of
families arranged in their series of most proximate affinities, the
first may present subdivisions, others, in irregular occurrence, may not
require them,—just as in the species of a genus, arranged also in the
series of their closest resemblances, one will present a stringent
adherence to the specific type, or all may do so, or all or some may
have a tendency to vary. Groupings of species are, however, of a less
natural character usually than are those of families, and generally are
artificial, being capriciously made to break down long genera, that the
required species may be more readily arrived at.
The characters which group families differ _inter se_. Thus in the Order
_Hymenoptera_, the family of the bees is essentially framed upon their
most distinguishing peculiarity—the tongue,—which in other families
becomes of secondary importance. In some the neuration of the wings,
their mode of folding, the form of the eyes, conjunctively with other
peculiarities of general structure, etc. etc., which point to the
differences in the economy that accompany all these, have successively
the same prominent position which the trophi take in the family of the
bees.
I have already recently alluded to the relations of affinity and
analogy, and it is desirable that some notion of the meaning and bearing
of these terms should be given, as, in the majority of modern works on
natural history, use is frequently made of them.
On carefully surveying any class or order of creatures, the mind
speedily becomes impressed by observing certain similitudes out of the
direct line of continuous connection, and therefore remote from the
strongest connecting links of positive relationship in the methodical
series. Induced thence to inspect them more closely, we presently
ascertain that what we at first conceived might be an error in their
collocation, arises from very strong resemblances in certain particular
features, but which are less important than those which directly unite
them, and may not be permitted to interrupt the order established. It
is, however, equally evident that they indicate relations which may not
be neglected.
Thus, although the succession be direct in the evolution of its primary
characteristics, the prominent features which so present themselves
establish the conviction of the existence of connections oblique to the
straight line, but all embraced within the normal conditions which bind
the group together. These are called relations of affinity. Pursuing
them, it is sometimes observed that nature, as it were, returns upon
itself, reproducing similar notes in another key.
These indications have led philosophical naturalists to surmise that the
true arrangement of natural objects is in groups, and not in a straight
and continuous line.
Several schemes have been suggested for the purpose of giving uniformity
to these groups, making them equivalents by associating together the
same numbers of allied forms, which again return in a circular series
upon themselves, and impinge upon other circles at the parallel points
of their circumference by affinities less direct than those which unite
them within their own circle.
Many novel views and interesting combinations have been thus elicited,
showing that very strong affinities lie in very divergent directions,
but no system has been hitherto devised which overrules the conflicting
difficulties that attend these arrangements. Whatever number may have
been adopted to bring nature within this circular system, it has always
been found that some, or several members, both in the circles
themselves, or in their series, is as yet deficient, and awaits either
discovery or creation.
The pursuit of such views stimulates profound investigation, and may
lead to valuable discoveries that will eventually give a loftier and
more philosophical character to the study of natural history than it has
hitherto possessed, and make it an attraction to the highest class of
mental powers. The key to the universe hangs at the girdle of the veiled
goddess; and happy the student who shall achieve possession of it, and
unlock the mysteries to the reverential gaze of mankind.
The _relation of analogy_ is different in _kind_, although the general
affinities which bind a class together are necessarily affinities in the
widest construction of the term; but the class being resolved into its
elements, those affinities, thus dissevered, no longer retain the
uniting links whereby the mass coheres. They, more correctly, stream
from their origin in parallelisms rather than in a continuous and
uninterrupted current; and these parallelisms present resemblances often
of a merely superficial character. As strong an instance as I can adduce
is possibly the analogical parallelism of the _Pentamera_ and the
_Heteromera_ in the _Coleoptera_, which are, however, bound by the
common affinity of being all beetles.
It is, nevertheless, often difficult to determine between the
relationships of affinity and analogy, for groups even in close
contiguity may also possess both. Thus, the normal _Ichneumones_ have
their analogues in the _Ichneumones adsciti_, if the comparison be
restricted to themselves, but these revert into the relationship of
affinity when a comparison is instituted between them and the adjacent
groups on the one side of the _Tenthredines_, or on the other of the
_Aculeata_, with which, when a relationship presents itself, it is
merely one of analogy. So, also, within the pentamerous _Coleoptera_ we
have a relationship of analogy between the _Staphylinidæ_ and the
_Histeridæ_, but it becomes one of affinity when it unites them within
this section of the class.
Innumerable other instances might be given readily, but these will
suffice to convey a notion of the relative meanings of the terms,
‘relation of affinity’ and ‘relation of analogy,’ which is all here
aimed at.
The problem naturalists have to solve is, “What is the natural system?”
We can clearly see that the systems adopted are not Nature’s, that they
are essentially imperfect, and that the science, even with all the force
of the intelligence that has been applied to it, is far from having
attained perfection. It still awaits the master mind that shall cope
with its difficulties, determine its intricacies, and, threading the
labyrinth, guide his enthusiastic disciples into the adytum of the
temple.
The subjects here brought under view admit of very considerable
development, and of strictly didactic and methodical treatment. It has
been my object only to gossip upon them, that I might stimulate
curiosity to undertake systematic study, by showing how interesting it
may become if earnestly pursued, being so fraught with instruction of
large compass.
Works on natural history have divers objects in view, and may be
intended either for popular and general distribution, or for special
scientific purposes, and in each case the mode of treatment will
materially differ. Many purposes may also be intended to be severally
met in the strictly and rigidly scientific treatment. They may be either
general methodical arrangements treated superficially, having no other
design than to give a sort of bird’s-eye view of the subject in its
wider distributions and broader landmarks, or they may treat of portions
of the large subject more specially; again, they may constitute
monographs of varying extent from a family to a genus; or they may
comprise loose descriptions of new species of old and well-established
genera; and some such, conjunctively with new species, establish
likewise new genera, indicating, at the same time, their proximate
position in the general series. The two latter classes are usually the
appendages to voyages and travels in distant unexplored countries, or
are the result of a careful collection of neglected tribes at home.
Each, thus, with its special application has its special construction;
but in the case of new species, I would strenuously counsel a full and
complete description, and urge as imperative the construction of a
specific character, formally framed to meet the condition of the
science, based upon the precise antecedents and existing state of the
genus to which such species belong.
Even assuming that the knowledge of species is the essential foundation
of the science, the preceding observations show that there is a higher
knowledge connected with the pursuit than this mere knowledge of
species, and yet from which it emanates. There is a higher object to be
achieved than the accumulation of a store of them, arranged in seemly
order, set with manifest taste, and named in accordance with the
accepted nomenclature. These are extremely pleasing to the eye, but the
intellect languishes over them in unsatisfied desire, craving more solid
aliment. There is besides room for observation on every side, either
confirmatory or original, and both are much needed, and must be
considerably augmented before it is accumulated in satisfactory
abundance; and until this be procured, existing systems can be viewed
merely as temporarily useful, for until all that nature can teach shall
be exhausted, perfection cannot be attained.
The many kinds of knowledge which the study subserves, and the
recreation and pleasure each affords, are a sufficient reply to the
sneering _Cui bono?_ of its detractors, who, when they urge that it
occupies time which might be more profitably employed, present
themselves but as the priests of the Fetish of the age, and may be told
that we use it only as a relaxation to necessary worldly toils. When
pursued, in cases where it can be so, in unmolested security, is there a
more salutary pursuit than that which inculcates the high veneration and
love which the study of nature should inspire towards the Great Parent
of all? What can compete with it in other studies? The investigation of
the works of the Almighty lead directly to the steps of the altar of
religion, and there we find the study of the Works confirmed by the
precepts of the Word, both inculcating humble reverence and fervent
love. Thus pursued, is it not a reply to every cavil?
------------------------------------------------------------------------
CHAPTER VII
BRIEF NOTICE OF THE SCIENTIFIC CULTIVATION OF BRITISH BEES.
WITH the great JOHN RAY dawns the scientific cultivation of British
bees. Before his time, the only entomological work which had been
published in England was Dr. Mouffett’s ‘Theatrum Insectorum.’ In this
work there is an ample account of the domestic bee, with gleanings from
many sources of some of its habits and economy, but there is no notice
of any insects, excepting some species of the genus _Bombus_, which may
be at all consorted with the social bee by affinities of structure or
identity of function.
In Ray’s correspondence with his disciples and friends, we have
straggling observations upon the habits of a few wild bees, especially
some jotted down by his diligent pupil, the distinguished Francis
Willughby. It is in Ray’s posthumous ‘Historia Insectorum,’ published in
1710, at the instance of the Royal Society, that we first find collected
together all that had been previously known of ‘British Bees.’ In that
work he describes them systematically. He there arranges the bees into
_Apis_ and _Bombylius_, which may be regarded almost as genera.
He divides _Apis_ into what may be considered as two sections, _Apis
domestica_ forming the first, and the second containing his _Apes
silvestres_, or wild bees. Nine of these are described and numbered
consecutively, which are followed by eleven descriptions unnumbered,
some of the latter having been supplied to him by Francis Willughby,
whose initials are attached to these, and amongst which we find the
description of the willow bee, subsequently, from this cause, named by
Kirby, from its original describer, and now universally known as
_Megachile Willughbiella_.
Ray’s second genus is _Bombylius_, identical, as far as it goes, with
the modern genus _Bombus_, excepting that it includes an _Anthophora_.
He here describes nineteen, all numbered. Ray’s names are phrases, the
mode of describing then prevalent in all the natural sciences, until the
happy introduction of the binomial system by the great genius of natural
history—LINNÆUS. These phrases are almost tantamount to the modern
specific character; but Ray unfortunately attaches no size, yet size
might have lent some aid to their modern determination.
Mr. Kirby was able to identify and introduce into his synonymy only a
few of Ray’s insects, from the defectiveness of the descriptions; the
following embrace all that could be verified:—
No. 1 of the _Apes silvestres_ is our _Anthidium mancatum_; No. 3, the
male of _Anthophora retusa_, the female of which being No. 4 of his
_Bombylii_; No. 4 of the _Apes_ is _Andrena nitida_: these comprise all
of those numbered which could be recognized. The first of the unnumbered
is the male of _Eucera longicornis_; the fourth is _Melecta punctata_;
the sixth is _Colletes fodiens_; the seventh is the male of _Osmia
bicornis_; and the ninth the celebrated _Megachile Willughbiella_.
In _Bombylius_ No. 1 is _Bombus lapidarius_; No. 2, _B. Raiellus_, named
by Mr. Kirby in honour of its great describer; No. 3 is _B. muscorum_;
No. 4 is the female of _Anthophora retusa_, as noticed above; No. 5 is
_Bombus terrestris_, as is also No. 6; No. 7 is the male of _B.
lapidarius_; No. 8 is _B. pratorum_; No. 9 is _B. sylvarum_; No. 10 is
_B. subinterruptus_; No. 11 is _B. hortorum_; No. 13 is _B.
Francillonellus_, and No. 17 is _Apathus Barbutellus_. Thus ten of the
_Apes silvestres_, and six of the _Bombylii_ are unidentified, and those
recognized may be placed correctly, by the aid I give in attaching Mr.
Kirby’s synonymy to the list of species added to each genus below.
Nothing of any moment thence intervened, until the Rev. W. Kirby, of
Barham, in Suffolk, made a careful and earnest collection of the
‘British Bees,’ with a view to their scientific description and
distribution. Stragglers were to be found in many entomological
cabinets, and some of their habits had been observed and recorded by
patient and attentive naturalists; but these collections were small,
very imperfect, and widely dispersed, until Mr. Kirby’s energy and
activity nurtured the idea, and carried it into execution, of bringing
into one focus the scattered notices and vagrant specimens he had seen
about.