CHAPTER XVIII.

PHYSIOLOGY OF THE CIRCULATORY ORGANS.

352. The walls of all the cavities of the heart are composed of

muscular fibres, which are endowed with the property of contracting

and relaxing, like the muscles of the extremities. The contraction and

relaxation of the muscular tissue of the heart, produce a diminution

and enlargement of both auricular and ventricular cavities. The

auricles contract and dilate simultaneously, and so do the ventricles;

yet the contraction and dilatation of the auricles do not alternate

with the contraction and dilatation of the ventricles, as the

dilatation of the one is not completed before the contraction of the

other commences. The dilatation of the ventricles is termed the

_di-as'to-le_ of the heart; their contraction, its _sys'to-le_.

353. The ventricles contract quicker and more forcibly than the

auricles, and they are three times longer in dilating than contracting.

The walls of the right ventricle, being thinner than the left, are more

distensible, and thus this cavity will contain a greater amount of

blood. This arrangement adapts it to the venous system, which is

more capacious than the arterial. The thicker and more powerful walls of

the left ventricle adapt it to expel the blood to a greater distance.

354. The valves in the heart permit the blood to flow from the

auricles to the ventricles, but prevent its reflowing. The valves at

the commencement of the aorta and pulmonary artery, permit the blood

to flow from the ventricles into these vessels, but prevent its

returning.

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352-366. _Give the physiology of the circulatory organs._ 352. What do

the contraction and relaxation of the muscular walls of the heart

produce? How do the auricles and ventricles contract and dilate? 353.

What is said of the contraction and dilatation of the ventricles in

the heart? How is the right ventricle adapted to its function? How the

left? 354. What is the use of the valves in the heart? Those of the

aorta and pulmonary artery?

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355. The function of the different parts of the heart will be given,

by aid of fig. 74. The blood passes from the right auricle (3) into

the right ventricle, (5,) and the tricuspid valves (6) prevent its

reflux; from the right ventricle the blood is forced into the

pulmonary artery, (7,) through which it passes to the lungs. The

semilunar valves (9) prevent this circulating fluid returning to the

ventricle. The blood, while passing over the air-cells in the lungs,

in the minute divisions of the pulmonary artery, is changed from a

bluish color to a bright red. It is then returned to the left auricle

of the heart by the pulmonary veins, (11, 11.)

[Illustration: Fig. 74. 1, The descending vena cava, (vein.) 2, The

ascending vena cava, (vein.) 3, The right auricle. 4, The opening between

the right auricle and the right ventricle. 5, The right ventricle. 6, The

tricuspid valves. 7, The pulmonary artery. 8, 8, The branches of the

pulmonary artery that pass to the right and left lung. 9, The semilunar

valves of the pulmonary artery. 10, The septum between the two ventricles

of the heart. 11, 11, The pulmonary veins. 12, The left auricle. 13, The

opening between the left auricle and ventricle. 14, The left ventricle.

15, The mitral valves. 16, 16, The aorta. 17, The semilunar valves of the

aorta.]

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355. Describe the course of the blood from the right auricle in the

heart to the lungs.

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_Observation._ If the blood is not changed in the lungs, it will not

flow to the pulmonary veins. This phenomenon is seen in instances of

death from drowning, strangling, carbonic acid, &c. The same is true,

but in a less degree, of individuals whose apparel is tight, as well

as of those who breathe impure air, or have diseased lungs.

356. The left auricle, (12,) by its contraction, forces the blood into

the left ventricle, (14.) The mitral valves (15) prevent its

reflowing. From the left ventricle the blood is forced into the aorta,

(16,) through which, and its subdivisions, it is distributed to every

part of the system. The semilunar valves (17) prevent its returning.

_Observation._ The parts of the circulatory organs most liable to

disease are the valves of the heart, particularly the mitral. When

these membranous folds become ossified or ruptured, the blood

regurgitates, and causes great distress in breathing. The operations

of the system are thus disturbed as the movements of the steam engine

would be if its valves were injured, or did not play freely.

357. The difference between the functions of the pulmonary artery and

aorta is, the former communicates with the right ventricle of the

heart, and distributes only impure blood to the lungs; the other

connects with the left ventricle of the heart, and distributes pure

blood to the whole body, the lungs not excepted. At the extremity of

the divisions of the aorta, as well as the pulmonary artery, are found

capillary vessels. This curious net-work of vessels connects with the

minute veins of the body, which return the blood to the heart.

_Observation._ The function of the veins of the systemic circulation

is similar to the office of the arteries in the lungs, and that the

veins of the pulmonic circulation transmit to the heart the pure, or

nutrient blood, and thus supply the arteries of the general system

with assimilating fluid.

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What is the effect when the blood is not changed in the lungs? 356.

Describe the circulation of the blood from the left auricle to the

general system. What part of the circulatory organs is most liable to

disease? What is the effect when the valves are diseased? 357. Give

the difference in the functions of the pulmonary artery and aorta.

Show the relation between the functions of the arteries and veins both

of the pulmonic and systemic circulation.

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358. The veins that receive the blood from all parts of the body,

follow nearly the same course as the arteries. The myriads of these

small vessels beneath the skin, and others that accompany the

arteries, at last unite and form two large trunks, called _ve'na ca'va

as-cend'ens_, and _de-scend'ens_.

_Observation._ A peculiarity is presented in the veins which come from

the stomach, spleen, pancreas, and intestines. After forming a large

trunk, they enter the liver, and ramify like the arteries, and in this

organ they again unite into a trunk, and enter the ascending vein, or

cava, near the heart. This is called the portal circulation.

359. The ventricles of the heart contract, or the "pulse" beats, about

seventy-five times every minute; in adults; in infants, more than a

hundred times every minute; in old persons, less than seventy-five

times every minute. The energy of the contraction of this organ varies

in different individuals of the same age. It is likewise modified by

the health and tone of the system. It is difficult to estimate the

muscular power of the heart; but, comparing it with other muscles, and

judging from the force with which blood is ejected from a severed

artery, it must be very great.

_Observation._ The phenomenon known under the name of pulse, is the

motion caused by the pressure of the blood against the coats of the

arteries at each contraction of the ventricles.

360. The following experiment will demonstrate that the blood flows

from the heart. Apply the fingers upon the artery at the wrist, at

two different points, about two inches apart; if the pressure be

moderately made, the "pulse" will be felt at both points. Let the

point nearest the heart be pressed firmly, and there will be no

pulsation at the lower point; but make strong pressure upon the lower

point only, and the pulsation will continue at the upper point,

proving that the blood flows from the heart, in the arteries, to

different parts of the system.

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358. What is the course of the veins? What peculiarity is observable

in the veins of the liver? 359. How often does the heart contract, or

the pulse beat, in adults? In infants? In old persons? What is said of

the energy of its contraction in different persons? How is the pulse

produced? 360. Demonstrate by experiment that the blood flows from the

heart.

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361. There are several influences, either separately or combined that

propel the blood from the heart through the arteries, among which may

be named,--1st. The contraction of the muscular walls of the heart.

2d. The contractile and elastic middle coat of the arteries aids the

heart in impelling the blood to the minute vessels of the system. 3d.

The peculiar action of the minute capillary vessels is considered, by

some physiologists, as a motive power in the arterial circulation.

4th. The pressure of the muscles upon the arteries, when in a state of

contraction, is a powerful agent, particularly when they are in active

exercise.

362. The following experiments will demonstrate that the blood from

every part of the system flows to the heart by the agency of the

veins. 1st. Press firmly on one of the veins upon the back of the

hand, carrying the pressure toward the fingers; for a moment, the vein

will disappear. On removing the pressure of the finger, it will

reappear, from the blood rushing in from below.

2d. If a tape be tied around the arm above the elbow, the veins below

will become larger and more prominent, and also a greater number will

be brought in view, while the veins above the tape are less distended.

At this time, apply the finger at the wrist, and the pulsation of the

arteries still continues, showing that the blood is constantly flowing

from the heart through the arteries, into the veins; and the

increased size of the veins shows that the pressure of the tape

prevents its flowing back to the heart.

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361. State the influences that propel the blood from the heart. 362.

Demonstrate by the first experiment that the blood flows to the heart.

By the second experiment.

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363. The influences that return the blood to the heart through the

veins, are not so easily understood as those that act on the blood in

the arteries. Some physiologists have imputed an active propulsive

power to the capillary vessels in carrying the blood through the

veins. This is not easily explained, and perhaps it is as difficult to

understand. An influence upon which others have dwelt, is the suction

power of the heart in active dilatation, acting as a _vis a fronte_

(power in front) in drawing blood to it.

364. Another influence that aids the venous circulation is attributed

to the propulsive power of the heart. It is not easy to comprehend how

this power of the heart can be extended through the capillary vessels

to the blood in the veins. Again, an important agency has been found,

by some physiologists, in the inspiratory movements, which are

supposed to draw the blood of the veins into the chest, in order to

supply the vacuum which is created there by the elevation of the ribs

and the descent of the diaphragm.

365. One of the most powerful causes which influence the venous

circulation, is the frequently-recurring action of the muscles upon

the venous trunks. When the muscles are contracted, they compress that

portion of the veins which lie beneath the swell, and thus force the

blood from one valve to the other, toward the heart. When they are

relaxed, the veins refill, and are compressed by the recurring action

of the muscles.

_Observation._ The physician, in opening a vein, relies on the

energetic contractions and sudden relaxations of the muscles, when he

directs the patient to clasp the head of a cane, or the arm of a

chair; these alternate motions of the muscles cause an increased flow

of blood to the veins of the ligated arm.

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363. What is said of the influences that return the blood to the

heart? What is said of the propulsive power of the capillaries? Of the

suction power of the heart? 364. Give another influence. State another

agency. 365. What is one of the most powerful causes which influence

venous circulation? Give practical observation.

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[Illustration: Fig. 75. An ideal view of the circulation in the lungs and

system. From the right ventricle of the heart, (2,) the dark, impure

blood is forced into the pulmonary artery, (3,) and its branches (4, 5)

carry the blood to the left and right lung. In the capillary vessels (6,

6) of the lungs, the blood becomes pure, or of a red color, and is

returned to the left auricle of the heart, (9,) by the veins, (7, 8.)

From the left auricle the pure blood passes into the left ventricle,

(10.) By a forcible contraction of the left ventricle of the heart, the

blood is thrown into the aorta, (11.) Its branches (12, 13, 13) carry the

pure blood to every organ or part of the body. The divisions and

subdivisions of the aorta terminate in capillary vessels, represented by

14, 14. In these hair-like vessels the blood becomes dark colored, and is

returned to the right auricle of the heart (1) by the vena cava

descendens, (15,) and vena cava ascendens, (16.) The tricuspid valves

(17) prevent the reflow of the blood from the right ventricle to the

right auricle. The semilunar valves (18) prevent the blood passing from

the pulmonary artery to the right ventricle. The mitral valves (19)

prevent the reflow of blood from the left ventricle to the left auricle.

The semilunar valves (20) prevent the reflow of blood from the aorta to

the left ventricle.]

366. The muscles exercise an agency in maintaining the venous

circulation at a point above what the heart could perform. As the

pulsations are diminished by rest, so they are accelerated by

exercise, and very much quickened by violent effort. There can be

little doubt that the increased rapidity of the return of blood

through the veins, is, of itself, a sufficient cause for the

accelerated movements of the heart during active exercise.

_Observation._ The quantity of blood in different individuals varies.

From twenty-five to thirty-five pounds may be considered an average

estimate in a healthy adult of medium size. The time in which the

blood courses through the body and returns to the heart, is different

in different individuals. Many writers on physiology unconditionally

limit the period to three minutes. It is undeniable that the size and

health of a person, the condition of the heart, lungs, and brain, the

quantity of the circulating fluid, the amount and character of the

inspired air, and the amount of muscular action, exert a modifying

influence. The time probably varies from three to eight minutes.

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366. What causes the accelerated movements of the heart during active

exercise?

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_Note._ Let the pupil review the anatomy and physiology of the

circulatory organs from fig. 75, or from anatomical outline plates,

No. 6 and 7.

A Treatise on Anatomy, Physiology, and Hygiene (Revised Edition) · 第 2 / 34 章