Relationships

The large genus _Coluber_ is much in need of revision. Its many

species, perhaps more than a score in all, occur in North America from

southern Canada south to Guatemala, in eastern and southwestern Asia,

in southern Europe, and in North Africa. All are active,

slender-bodied snakes having smooth scales in few rows, and having

large eyes with well developed vision. The North American species fall

into two natural groups, the typical racers, and the whip snakes,

often assigned to a separate genus, _Masticophis_ (Ortenburger, 1928).

The whip snakes are more specialized than the typical racers in having

the eyes more enlarged, and the body form more slender and attenuate,

with number of scale rows more reduced. The racers of the Old World

are more diverse. Inger and Clark (1943) suggested a partitioning of

the genus _Coluber_ on the basis of the pattern by which scale rows

are reduced, from the maximum number on the forebody to the minimum

number at the posterior end of the body, supplemented by certain

characters of the hemipenis and of the cephalic scutellation. Besides

_Coluber_ and _Masticophis_ these authors recognized within the group

the genus _Platyceps_ with several species in southern Europe and

southwestern Asia; _Zamenis_ with several species in the same region

and in North Africa, and _Haemorrhois_, a monotypic genus of Spain,

North Africa and several Mediterranean islands. Although apparently

valid in principle, this arrangement has not been generally followed.

Exclusive of those species groups whose assignment to the genus

_Coluber_ are somewhat questionable, the remaining species in the

genus are: _C. constrictor_ occurring throughout most of the United

States and south along a narrow Atlantic coastal strip of Mexico to

Guatemala; _C. oaxacae_ of southern Mexico; and _C. spinalis_ of

northern China. _C. oaxacae_ is poorly known as only a few specimens

have been collected, but seemingly it is a near relative and

derivative of _C. constrictor_, especially of that species'

southernmost population. _C. spinalis_ is much more distinct, as might

be expected from its geographical remoteness. It is a slender, active

snake, of olive coloration dorsally with 17 scale rows and a bright

yellow, black-edged dorsal stripe and yellow ventral surface. It is

relatively small (up to 755 millimeters snout-vent length) and is

partial to riparian habitats but is also found in forests and in dry

and barren regions (Pope, 1935:224-226). It is known to feed upon

lizards.

Natural History of the Racer Coluber constrictor · 第 28 / 47 题