Creek, 7 mi. SW Daisy. _Caddo_: ANSP 100, Washita River, Fort Cobb.
_Choctaw_: OU 27126, Mayhew Creek, 2 mi. NW Boswell. _Comanche_: OU
4130, 4266, 5390, 8333, 12953, 19986, Wichita Mountains Wildlife
Refuge. _Jackson_: OU 13012, 6 mi. E El Dorado. _Kiowa_: CNHM 15474.
_Le Flore_: OU 6791, Kiamichi River, 8 mi. W Arkansas State Line.
_McCurtain_: OU 2149-50, 2152, 2155, 17126-28, 17185, 2 mi. SW
Smithville; USNM 70397, Red River. _Marshall_: KU 40175-76, 50830-31,
50847, OU 27290, 27297, 27562-63, TU 16076 (5), 16175 (6), 16662 (5),
Lake Texoma, 2 mi. E Willis; KU 50832, mouth of Caney Creek, 4 mi. SW
Kingston. _Pushmataha_: OU 2151, 2157; OU 11365, Buffalo Creek, 5 mi.
NW Tuskahoma.
TEXAS: _Archer_: TU 16174, 16668-69, Lake Diversion. _Bell_: SM
5667-69, Nolan Creek. _Bosque_: KU 47121, 7 mi. below Whitney Dam,
Brazos River. _Brazos_: BCB 4436, 10 mi. E College Station; BCB 4437,
17 mi. S College Station; BCB 4438, 4 mi. N Bryan; KU 50833, 4 mi. W
College Station; SM 2556, TCWC 472, Wickson Lake; TCWC 539, Little
Brazos River; TCWC 4692, 8 mi. NE Bryan; TCWC 5121, 2 mi. S College
Station; TCWC no number. _Clay_: TCWC 7258, 8 mi. NW Ringgold,
Montague County; TU 16667.1, 3 mi. W Byers. _Dallas_: MCZ 3987, "near"
Dallas; ANSP 13243, Dallas. _Donley_: ANSP 13440, S of Clarendon.
_Eastland_: KU 3132, Cisco. _Galveston_: TCWC 7251, Alta Loma.
_Harris_: UMMZ 92753, Little Cypress Creek, 1 mi. N Westfield; USNM
94335-36, "near" Houston. _Harrison_: USNM 95386, 16.5 mi. SE Caddo
Lake. _Hill_: TU 14169, Richland Creek, 0.7 mi. W Mertens. _Leon_:
CNHM 46290, 5 mi. W Marquez; TCWC 8994, 8996, 6 mi. NW Normangee.
_Liberty_: TU 14402, 14417, Trinity River, "near" jct. with Big Creek.
_McLennan_: BCB 4665-66, 6 mi. NNE McGregor; SM no number, 2037, 2452,
2552, 2558, 2560, 2640, 5263, 6533, Lake Waco; SM 0185, Middle Bosque
River; SM 2104, 6732, Upper Bosque River; SM 5072, Bull Hide Creek; UI
2399, 1.5 mi. W China Springs; UMMZ 64063, Waco; USNM 55601.
_Madison_: TCWC 471, 517, Twin Lakes. _Montgomery_: TCWC 540, 3 mi. S
Conroe. _Nacogdoches_: TNHC 14112, Legg Creek, 5 mi. S Douglass.
_Orange_: UMMZ 117060, 3 mi. S Orange; USNM 94456-57, Orange.
_Randall_: TTC 576, Palo Duro Canyon, 15 mi. SE Canyon. _Shackelford_:
TU 14547, Clear Fork Brazos River, Fort Griffin State Park.
_Somervell_: TCWC 8995, TU 14559 (4), Brazos River, 5-6 mi. E Glen
Rose. _Trinity_: SM 2889, Groveton. _Walker_: TNHC 20829, 5 mi. E New
Waverly. _Waller_: TNHC 14068, 2.7 mi. E Brazos River on US 90.
_Williamson_: MCZ 1627 (2); TU 14348, San Gabriel River, 6.5 mi. E
Georgetown. _County unknown_: ANSP 13448, Wichita River; USNM 7640,
Brazos River.
_Records in the literature._--LOUISIANA: _Cameron_: Sabine Refuge
(Cagle and Chaney, 1950:386).
OKLAHOMA: _Le Flore_: 6 mi. W Page. _McCurtain_: 14 mi. SE Broken Bow
(Trowbridge, 1937:301).
TEXAS: _Bosque_: Bosque River, "near" Valley Mills (Strecker, 1928:6).
_Harris_: Addicks (Brown, 1950:250). _Henderson_: Cedar Creek
(Strecker, 1926a:7). _Jefferson_: 12 mi. SW Port Arthur (Guidry,
1953:56). _Liberty_: Daisetta (Brown, _loc. cit._); San Jacinto River
(Strecker, 1915:15). _McLennan_: "near" Crawford (Brown, _loc. cit._).
_Orange_: 1 mi. N Bridge City (Guidry, _loc. cit._). _Tarrant_:
Trinity River, Fort Worth (Stejneger, 1944:66). _Taylor_: Abilene
(KKA). _Tyler_: Colmisneil (Siebenrock, 1909:603). _Walker_: 6 mi. E
Huntsville (TCWC 329, listed in card file). _Wheeler_: 5 mi. N Wheeler
(Brown, _loc cit._).
=Trionyx ater= Webb and Legler
Black Softshell
_Trionyx ater_ Webb and Legler, Univ. Kansas Sci. Bull., 40:21,
pls. 1 and 2, 1960, April 20.
_Type._--Holotype, KU 46903, alcoholic female; obtained 16 km. S
Cuatro Ciénegas, Coahuila, México, by John M. Legler (and party),
September 6, 1958.
_Range._--Basin of Cuatro Ciénegas, central Coahuila, Mexico (see map,
Fig. 22).
_Diagnosis._--Posterior margin of carapace of some females having fine
corrugations, edge often ragged, and no pale outer margin; septal
ridges reduced in adult males; over-all dorsal coloration (in
preservative) dark, lacking contrasting patterns.
_Description._--Plastral length of adult male, 9.6 centimeters (KU
46911); of largest female, 18.4 centimeters (KU 46903).
Adult male: anterior edge of carapace smooth; septal ridges reduced;
pale outer rim, and small, whitish, dots posteriorly on carapace;
surface of carapace slightly gritty or sandpapery posteriorly; snout
broadened; over-all dorsal coloration dark gray or slate; contrasting
pattern on soft parts of body lacking; ventral surface whitish having
few blackish marks posteriorly on undersurface of carapace.
Females: posterior margin of carapace usually having fine
corrugations; edge of carapace posteriorly often ragged; pale rim of
carapace absent; mottled and blotched pattern not contrasting on
blackish carapace; dorsal surface of soft parts of body dark gray or
slate, lacking contrasting pattern; ventral surface of carapace and
posterior part of plastron usually having many blackish flecks and
markings; tubercles lacking on anterior edge and in center of carapace
posteriorly; septal ridges well developed.
Medial angle of epiplastron (as observed through overlying skin) bent
at angle of approximately 90 degrees. Other osteological characters
presumably as in _spinifer_.
Range in length of plastron (cm.) of 11 females (mean follows
extremes); 10.8-18.4, 15.0; proportional measurements of 12 specimens
(including adult male, mean follows extremes): PL/HW, 4.70-5.43, 4.93;
CL/CW, 1.28-1.43, 1.32; CL/PCW, 1.98-2.42, 2.15; HW/SL, 1.22-1.58,
1.37; CL/PL, 1.29-1.44, 1.36; some females (especially KU 46908) have
noticeably elongate carapaces.
_Variation._--Corrugations best developed on two largest females (KU
46903, 46906), even present on ventral surface of carapace posteriorly
and on dorsal surface of tail; development of corrugations not
ontogenetic phenomenon as posterior margin relatively smooth on KU
46908 (plastral length, 16.0 cm.) but relatively rugose on KU 46909,
which is smaller (plastral length, 13.9 cm.); smallest female (KU
46904) and adult male having posterior margin smooth; smallest female
having indication of pale outer rim and small whitish dots posteriorly
on carapace, and dark, obtusely-angular line, connecting anterior
margins of orbits; blackish marks on ventral surface reduced on KU
46904, 46910, 46912, and UI 43510; UI 43510 (plastral length, 16.3
cm.) resembles _T. s. emoryi_ in having more contrasting mottled
pattern on carapace and limbs, indication of pale outer rim on
carapace, and dark line connecting anterior margins of orbits; ventral
surface of tail and hind limbs often tinged with red.
Color notes from life of young female, topotype (KU 53755) are:
mottled carapace dark brown, pale areas buff; dorsal surface of head
mottled, olive-brown, pale areas buff; iris orange-buff; upper and
lower lips yellow-orange; dorsal surface of limbs olive-brown having
yellow to buff suffusion and small blackish marks; pale areas on
webbing yellow; ventral surface whitish having yellow at margin of
carapace, on neck and limbs.
_Comparisons._--_T. ater_ most closely resembles _T. spinifer_
(especially the subspecies _emoryi_) in having a gritty or
"sandpapery" carapace (reduced, tubercles more scattered), whitish
dots on posterior third of carapace (small females and adult male) and
a dark line connecting anterior margins of orbits (smallest female).
Prior to acquiring the characteristic darkened, dorsal ground color,
the pattern on the head and limbs seems to be that of _T. s. emoryi_.
_T. ater_ resembles _T. muticus_ in having reduced septal ridges in
males, a smooth anterior edge of carapace (especially males), and no
enlarged prominences on the anterior edge of the carapace or
posteriorly in the center of the carapace on large females. _T. ater_
resembles _T. ferox_ in having an over-all dark coloration dorsally
with no contrasting patterns on adults.
_T. ater_ probably is a small species resembling _T. muticus_ and _T.
spinifer emoryi_. The head is wide in _T. ater_, resembling that of
_T. ferox_, and closely approaching that of _T. spinifer emoryi_ and
_T. s. guadalupensis_. _T. ater_ resembles _T. ferox_ and _T. s.
emoryi_ in having a narrow carapace. _T. ater_ resembles _T. s.
emoryi_, _T. s. guadalupensis_ and _T. s. pallidus_, but differs from
_T. muticus_, _T. ferox_ and the other subspecies of _T. spinifer_ in
having the carapace widest farther posterior than one-half the length
of the carapace. _T. ater_ resembles _T. ferox_ and _T. s. emoryi_ in
shortness of snout. The plastron is short in _T. ater_ and most
closely resembles that of _T. s. pallidus_, _T. s. guadalupensis_, and
_T. s. emoryi_.
_Remarks._--_T. ater_ is confined to permanent, clear-water ponds in
the basin of Cuatro Ciénegas. The male and 11 females (KU) were taken
at the type locality (a pond known locally as Tío Candido); the other
female (UI 43510) was taken from a pond approximately seven miles
northward (known locally as Anteojo). _T. spinifer emoryi_ also occurs
in the basin of Cuatro Ciénegas. Males and females of _emoryi_ were
collected in the Río Mesquites (Río Salado drainage) that drains the
basin; two adult males of _emoryi_ were taken from the clear-water
ponds--one from the type locality of _ater_ (KU 46907), and the other
(KU 53757) from a pond (known locally as El Mojarral) from which no
_ater_ were obtained. This demonstrated sympatry indicates that the
two kinds are not conspecific.
However, the nature and frequency of occurrence of characters of _T.
ater_, suggest that it is subspecifically related to _T. spinifer_--in
effect, a darkened race of _T. s. emoryi_. The diagnostic characters
of fine corrugations on the posterior margin of the carapace and
blackish marks on the ventral surface do not occur on every female of
_ater_. Too, the dorsal coloration of living females (dark brown-buff)
is paler than that of preserved specimens (dark gray-slate).
Furthermore, a hatchling (CNHM 47367) recorded from Cuatro Ciénegas,
Anteojo, is not distinguishable from _emoryi_.
The mention of absence of septal ridges in males of _T. ater_ in the
original description (Webb and Legler, 1960:22) should be amended. The
septal ridges in the only known adult male are reduced; a small,
whitish ridge is present on the medial surface of each nostril, but is
not conspicuous in anterior view. The one adult male of _ater_ is
distinguished from _T. s. emoryi_ principally on the over-all dark,
dorsal coloration with concomitant loss of pattern, the noticeably
broadened snout, and the reduced septal ridges. The last character
mentioned possibly is variable in _ater_ (and in _emoryi_ in this
region) in view of the variation in development of the ridge on four
male _emoryi_ from the basin: well-developed on KU 53757 (Mojarral)
and KU 46907 (Tío Candido); reduced on KU 53752 (Río Mesquites),
resembling development in _ater_; and, reduced on right side only on
KU 53753 (Río Mesquites).
Presumably, the continued erosive action at the headwaters of the Río
Salado has permitted the invasion of this drainage into the formerly
isolated basin of Cuatro Ciénegas. In the basin, however, I know of no
evidence of a direct aquatic contact between the headwater streams and
the isolated, clear-water, ponds. How _emoryi_ entered the ponds is
unknown. Some of the ponds are tapped by small, man-made, irrigation
canals, but, so far as I know, these are not connected to the river.
The ponds have permanent water and are often separated by several
miles of arid environment. Overland dispersal between waterways is
possible in time of flooding. Local residents tell of the infrequent
sale of softshells in Cuatro Ciénegas, which hints at their dispersal
via the agency of man. The underlying gypsum substrate of the valley
has been subjected to considerable erosion; the ponds observed have
deep holes, and small caverns and grottos. There are conflicting
reports concerning subterranean connections between ponds. Possibly
there are underwater connections between some ponds and the headwater
streams of the Río Mesquites. Whatever the dispersal route for
_emoryi_ into the ponds has been, it is strange that the same route
has not been traversed by _ater_, permitting its occurrence in the Río
Mesquites.
On the basis of morphological criteria, I suspect that _ater_ and
_emoryi_ are genetically compatible. Possibly there is only sporadic
entrance of _emoryi_ into the ponds inhabited by _ater_, or the
accessible dispersal routes for _emoryi_ have been relatively recent
and there has been insufficient time for genetic adaptation. _T. ater_
is maintained as a full species because of the occurrence of two
distinct males (KU 46907, _emoryi_, and KU 46911, _ater_) in the same
pond (Tío Candido, the type locality). These two specimens are
contrasted in a photograph accompanying the type description (Webb and
Legler, 1960: Pl. II). The restricted distribution of _ater_, and its
characteristics suggest a relict population derived from a
_ferox_-like ancestor that may be in the process of becoming extinct.
There are two specimens in the CNHM recorded from Cuatro Ciénegas. One
is a female (CNHM 55661) having a plastral length of 19.0 centimeters,
and no specific locality other than Cuatro Ciénegas. I examined this
specimen before I knew of the existence of _ater_, and noted no
unusual features; I have not re-examined the specimen. It is
considered representative of _emoryi_. The second is a hatchling (CNHM
47367) having a plastral length of 3.2 centimeters, recorded from
Cuatro Ciénegas, Anteojo. The carapace is dark tan having small
whitish dots intermixed with a few indistinct, small, blackish specks
posteriorly. The specimen is indistinguishable from _emoryi_.
_Specimens examined._--Total 12, as follows: COAHUILA: KU 46903-06,
46908-12, 53755-56, 16 km. S Cuatro Ciénegas; UI 73510, 5.7 mi. W
Cuatro Ciénegas.
_Records in the literature._--Schmidt and Owens (1944:103) record
_emoryi_ from Cuatro Ciénegas (no museum numbers listed); presumably
their reference is to CNHM 55661.
=Trionyx muticus= Lesueur
Smooth Softshell
_Range._--United States from extreme western Pennsylvania, southern
Minnesota and South Dakota south to the Gulf of Mexico in Alabama, the
western end of the panhandle of Florida, and the eastern half of Texas
(see map, Fig. 22.)
[Illustration: FIG. 22. Geographic distribution of _Trionyx ater_
and _Trionyx muticus_. 1. _T. muticus muticus_. 2. _T. muticus
calvatus_. 3. _T. ater_.]
_Diagnosis._--Septal ridges absent; anterior edge of carapace smooth,
lacking prominences; juvenal pattern of large dusky spots (sometimes
ocellate), or small dusky (not black), dots and short lines; side of
head usually devoid of markings except for pale, usually
uninterrupted, postocular stripe.
Size small; head narrow; snout long; ventral surface of supraoccipital
spine broad proximally, lacking median ridge; foramen magnum evenly
rounded, ovoid; opisthotic-exoccipital spur absent; distal part of
opisthotic wing truncate; lateral condyle of articular surface of
quadrate tapered posteriorly, smaller than medial articular surface;
angle of epiplastron obtuse, approximately 100 degrees; callosity on
epiplastron sometimes covering entire surface; bony bridge wide in
relation to length.
_Description._--Septal ridges absent; external characteristics
variable (see accounts of subspecies); range in length, in
centimeters, of plastron of ten largest specimens of each sex, (mean
follows extremes), males, 11.8-14.0, 12.3; females, 17.7-21.5, 18.9;
ontogenetic variation in PL/HW, mean PL/HW of specimens having
plastral lengths 7.0 centimeters or less, 4.16, ranging from 7.1 to
13.0 centimeters, 5.82, and, exceeding 13.0 centimeters, 7.04; little
ontogenetic variation in CL/CW, mean CL/CW of specimens having
plastral lengths 8.0 centimeters or less, 1.15, and exceeding 8.0
centimeters, 1.16; mean CL/PCW, 1.97; mean HW/SL, 1.22; mean CL/PL,
1.39.
Greatest width of skull usually at level of squamosal (79%); foramen
magnum ovoid; opisthotic-exoccipital spur usually absent (97%); distal
part of opisthotic wing truncate, sometimes visible in dorsal view;
lateral condyle of articular surface of quadrate tapered posteriorly,
smaller than medial articular surface; maxillaries not in contact
above premaxillaries; combination of seven neurals, seven pairs of
pleurals, and contact of seventh pair of pleurals (38%), or eight
neurals, seven pairs of pleurals, and separation of seventh pair of
pleurals (41%); angle of epiplastron obtuse, greater than 90 degrees;
callosities well-developed, frequently on preplastra and epiplastron
of adults.
_Comparisons._--The absence of septal ridges distinguishes _muticus_
from _ferox_, all subspecies of _spinifer_, and _ater_ (ridges are
reduced in males of _ater_). The smooth anterior edge of the carapace
distinguishes _muticus_ from all other American kinds except _ater_
and some individuals of _T. s. emoryi_. _T. muticus_ resembles only
_ater_ and _ferox_ in usually lacking a well-defined, contrasting
pattern of blackish marks on the dorsal surface of the limbs. _T.
muticus_ resembles _ferox_ and differs from _spinifer_ and _ater_ in
lacking a gritty or "sandpapery" carapace on adult males. Adult males
of _T. muticus calvatus_ and some individuals of _T. m. muticus_ from
the Colorado River in Texas further resemble _ferox_ in having
postocular stripes with thick black borders.
_T. muticus_ is the smallest species in North America; the maximum
size of the plastron in adult males is approximately 14.0 centimeters
(16.0 cm. in _spinifer_) and of adult females 21.5 centimeters (31.0
cm. in _spinifer_). Males and females of _muticus_ are sexually mature
at approximately the same size as some _T. s. emoryi_; also, the great
development of the plastral callosities in _muticus_ corresponds to
that in some _emoryi_. The head is narrower in _muticus_ than in
_ferox_ or _spinifer_. The carapaces of specimens of _muticus_
exceeding plastral lengths of 8.0 centimeters are wider than those of
_ferox_, _ater_, _T. s. emoryi_ and _T. s. guadalupensis_ of
corresponding size. _T. muticus_ differs from _ater_ and three
subspecies of _spinifer_ (_pallidus_, _guadalupensis_, _emoryi_) in
having the carapace widest at a plane approximately one-half the
length of the carapace. The snout is longer in _muticus_ than in
_ferox_ and _spinifer_. _T. muticus_ differs from _ferox_ but
resembles _spinifer_ in having a relatively short plastron.
The skulls of _muticus_ differ from those of _ferox_ but resemble
those of _spinifer_ in usually having the skull widest at the level of
the squamosals. Skulls of _muticus_ resemble those of _ferox_ but
differ from those of _spinifer_ in usually lacking a well-developed
opisthotic-exoccipital spur. Skulls of _muticus_ are different from
those of _ferox_ and _spinifer_ in having the 1) ventral surface of
the supraoccipital spine widest proximally, lacking a medial ridge, 2)
foramen magnum ovoid, 3) distal part of opisthotic wing truncate, 4)
lateral condyle of articular surface of quadrate tapered posteriorly,
smaller than medial articular surface, and 5) maxillaries not in
contact above premaxillaries.
Plastrons of _muticus_ differ from those of _spinifer_ and _ferox_ in
having an obtusely-angled epiplastron, relatively large callosities
in adults, and a wide hyo-hypoplastral bridge (in relation to length).
_Remarks._--Agassiz (1857:399) regarded Lesueur's _Trionyx muticus_ as
the type species of the genus _Amyda_ and the only species known to
belong to the genus _Amyda_. Stejneger (1944:7, 9, 12) proposed the
generic name _Euamyda_ as a new name for the North American _Amyda
mutica_ as understood by Agassiz. _Euamyda_ was proposed for use only
if Agassiz's understanding was found to be correct. Actually,
Stejneger thought that the Old World and New World kinds concerned
were congeneric, and that the type species of the genus _Amyda_ was
the Old World species _Amyda javanica_ Schweigger (= _Testudo
cartilaginea_ Boddaert).
If _Trionyx muticus_ Lesueur is considered to be generically distinct
from other soft-shelled turtles, _Euamyda_ Stejneger, 1944, is
available as a generic name with _Trionyx muticus_ Lesueur, 1827, as
the type species (by monotypy).
_Geographical variation._--_Trionyx muticus_ shows no obvious
character gradients; the variation is mostly discontinuous and unlike
that in _T. spinifer_. On the basis of differences in the juvenal
pattern and pattern on head, _T. muticus_ can be divided into two
subspecies.
=Trionyx muticus muticus= Lesueur
Midland Smooth Softshell
Plates 45, 46, and 53
_Trionyx muticus_ Lesueur, Mém. Mus. Hist. Nat. Paris, 15:263,
pl.7, December, 1827.
_Trionyx muticus muticus_ Webb, Publ. Mus. Nat. Hist. Univ. Kansas,
11:520, August 14, 1959.
_Potamochelys? microcephala_ Gray, Proc. Zool. Soc. London, p. 87,
1864.
_Type._--Lectotype, Museum d'Histoire Naturelle, Paris, No. 8813;
dried carapace and plastron; obtained from the Wabash River, New
Harmony, Posey County, Indiana, by C. A. Lesueur in August, 1827 (Pl.
53).
_Range._--Central United States; in the Mississippi River drainage
from extreme western Pennsylvania, southern Minnesota and South Dakota
south to Tennessee, Louisiana and Oklahoma; streams of the Gulf Coast
drainage from the Mississippi River in Louisiana westward into Texas
including the Colorado River drainage (see map, Fig. 22).
_Diagnosis._--Juvenal pattern of dusky dots and usually short lines or
bacilliform marks; ill-defined pale stripes on snout usually evident
just in front of eyes; pale postocular stripe lacking thick, black
borders that are approximately one-half width of pale stripe (except
some in the Colorado River drainage of Texas).
_Description._--Plastral length of smallest hatchling, 2.1 centimeters
(INHS 3458); of largest male, 14.0 centimeters (CNHM 92003); of
largest female, 21.5 centimeters (KU 2308).
Juvenal pattern of dusky, grayish marks lacking sharp margins, and
usually consisting of both small spots and short streaks or dashes,
the former predominating; short streaks or dashes occasionally lacking
(TU 14375, Pl. 45, bottom, left; UMMZ 92751); markings variable in
number, few and widely spaced, or several and closely approximated
(Pl. 45, top, topotypes); pale rim separated from ground color by
ill-defined, dusky margin; pattern on adult males well-defined
resembling that of hatchlings (TU 16172.1, 16173), scarcely
discernable (TU 13294), or absent (TU 1242); mottled and blotched
pattern on carapace usually contrasting in large females.
Pale stripes extending forward from eyes usually not more than half
distance to tip of snout; inner borders of pale stripes on snout
usually absent or dusky and indistinct, occasionally blackish (TU
14606); outer borders of pale stripes darker than inner borders,
usually blackish; pale stripes on snout occasionally absent (CNHM
7845, UMMZ 92665, TU 5989, none of these specimens being large
females); pale postocular stripe having narrow, dusky or blackish
borders (especially UMMZ 92751, TU 14436); pale postocular stripe
usually complete, occasionally interrupted having prominent
dark-bordered anterior segment just behind eye (TU 14416); lower
border of postocular stripe usually in contact with dusky postlabial
line; no other markings on side of head; pattern on dorsal surface of
soft parts of body not contrasting, composed of closely approximated
fine markings that are little darker than background, over-all
coloration pale grayish; occasionally, few larger and more contrasting
markings on hind limbs (UMMZ 92751, TU 14436).