_Records in the literature._--ONTARIO: _Carleton_: Ottawa

(questionable record). _Essex_: Point Pelee. _Haldimand_: Dunville.

_Kent_: Lake St. Clair. _Norfolk_: Long Point. _Oxford_: Beachville.

_Wentworth_: Hamilton Bay (Logier and Toner, 1955:51).

QUEBEC: _Iberville_: Richelieu River at Iberville (Logier and Toner,

1955:51).

ALABAMA: _Lawrence_: Courtland (Stejneger, 1944:53).

ILLINOIS: _Boone_: Belvidere. _Bureau_: Bureau. _Cass_: Chandlerville.

_Clay_: Louisville (Cahn, 1937:189). _Cook_: Lake Michigan (Kennicott

_in_ Stejneger, 1944:44); Evanston (Necker, 1939:10); Chicago (Schmidt

and Necker, 1935:76). _Crawford_: Robinson. _Douglas_: northern part

of county (P. W. Smith, 1947:39). _Fayette_: Vandalia. _Fulton_:

Ellisville (Cahn, _loc. cit._). _Grundy_: Morris (Stille and Edgren,

1948:201). _Jackson_: Jacob (Cagle, 1942:158). _Jersey_: Grafton

(Cahn, _loc. cit._). _Kane_: Batavia; Dundee Game Farm (Stille and

Edgren, _loc. cit._). _Kankakee_: Kankakee River near Altort (Necker,

_loc. cit._). _Lake_: Fox Lake. _LaSalle_: Streator (Cahn, _loc.

cit._). _Lawrence_: (Hahn _in_ Stejneger, 1944:44). _Lee_: symbol on

map (Cahn, _loc. cit._). _McHenry_: McHenry (Stille and Edgren, _loc.

cit._). _Macon_: Decatur. _Macoupin_: Carlinville (Cahn, _loc. cit._).

_Ogle_: Oregon (Garman _in_ Cahn, _loc. cit._). _Randolph_: Chester,

Reily Lake. _Rock Island_: Barstow, Hillsdale, Rock Island (Cahn,

_loc. cit._). _Saline_: Horseshoe Lake (Stein, 1954:312).

_Stephenson_: Freeport (Cahn, _loc. cit._). _Union_: Bluff Lake

(Garman _in_ Cahn, _loc. cit._). _Whiteside_: Sterling, symbol on map

(Cahn, _loc. cit._). _Williamson_: Marion (Cagle, 1942:158).

_Winnebago_: Rockton; symbol in western part of county (Cahn, _loc.

cit._). _County unknown_: Fox River (Yarrow, 1882:29).

INDIANA: _Brown_: 1 mi. below Helmsburg (Myers, 1927:339). _Clay_: Eel

River (Kirsch _in_ Stejneger, 1944:45). _Franklin_: (Hughes _in_

Stejneger, _loc. cit._). _Jasper_: Jasper-Pulaski Game Preserve

(Swanson, 1939:690). _Jefferson_: Madison (Myers, _loc. cit._).

_Marion_: Irvington (Stejneger, _op. cit._:55). _Marshall_: 2 mi. NW

Culver (KKA). _Monroe_: Bloomington (McLain _in_ Stejneger, _op.

cit._:45). _Newton_: Lake Village (Stille and Edgren, _loc. cit._).

_Posey_: Wabash River at New Harmony (Lesueur, 1827:257). _Starke_:

Grant (Stille and Edgren, _loc. cit._). _Steuben_: Fish Creek "near"

Hamilton (Stejneger, _op. cit._:53). _County unknown_ (Knox or

Starke): USNM 72387, Knox (Stejneger, _op. cit._:55); "White Water

valley," east-central part of state (Butler, 1894:224). USNM 8359 (=

_Trionyx spinifer asper_) has been erroneously recorded from Madison,

Indiana, by Yarrow (1882:29) and Hay (1892:145); see discussion by

Cahn (1937:200) and Stejneger, (_op. cit._:73, 75).

KENTUCKY: _Edmonson_: Green River, Mammoth Cave National Park

(Hibbard, 1936:281). _Fleming_: Fox Creek (Welter and Carr, 1939:130).

_Jefferson_: (Funkhouser, 1925:71). _Morgan_: (Stejneger, 1944:54).

_County unknown_: Ohio and Pond rivers (Funkhouser, _loc. cit._).

MICHIGAN: _Berrien_: mouth of St. Joseph River at St. Joseph (Lagler,

1943:303). _Eaton_: Brookfield; Olivet (Clark _in_ Ruthven, Thompson

and Thompson, 1912:133). _Genesee_: (Miles _in_ Ruthven, Thompson and

Thompson, _loc. cit._). _Iosco_: (Lagler, 1943:283, symbol on map).

_Kent_: (Lagler, _loc. cit._). _Montcalm_: (Clark _in_ Ruthven,

Thompson and Thompson, _loc. cit._). _Muskegon_: Muskegon River "near"

Muskegon (Lagler, _op. cit._:303). _Van Buren_: Reynolds Lake, 2.5 mi.

E Lawrence (Edgren, 1942:180).

MISSISSIPPI: _De Soto_: Lake Cormorant (Stejneger, 1944:55). _Holmes_:

Thornton (Cook, 1946:185). _Humphreys_: Belzoni (Stejneger, _loc.

cit._). _Sunflower_: _Warren_: Vicksburg, Eagle Lake (Cook, _loc.

cit._). _Washington_: Lake Washington (Smith and List, 1955:125);

Greenville (Stejneger, _loc. cit._).

NEW YORK: _Albany_: Hudson River at Albany (DeKay, 1842:7); Mohawk

River at Cohoes (Eights _in_ Bishop, 1923:120). _Cattaraugus_:

Allegheny River and Red House Lake in Allegheny State Park (Eaton,

1945:115). _Chautauqua_: Lake Chautauqua (DeKay, _loc. cit._).

_Monroe_: Braddocks Bay and Long Pond on Lake Ontario (Wright,

1919:8). _Saratoga_: Hudson River near Baker's Falls (restricted

locality, Rafinesque, 1832:64). _County unknown_: Lake Cayuga; Mohawk

River (DeKay, _loc. cit._).

OHIO (Conant, 1951:158-59, 264, except records from Allen, Geauga and

Noble counties): _Allen_: Sugar Creek, 6 mi. N Lima (Adler and Dennis,

1960:27). _Ashland_: Long Lake, Lake Twp.; Black Fork, Sec. 27, Green

Twp. _Athens_: Hocking River "near" Athens; "near" Fisher, Alexander

Twp. _Auglaize_: Pusheta Creek, west of Wapakoneta. _Brown_: White Oak

Creek, 1 mi. N Higginsport. _Butler_: Oxford. _Champaign_: Mad River,

4 mi. SW Urbana. _Coshocton_: Walhouding River, below dam. _Defiance_:

Auglaize River, Shawnee Scout Camp, Defiance Twp. _Erie_: Huron;

Sandusky. _Fairfield_: Buckeye Lake. _Franklin_: Alum Creek,

Westerville; Columbus. _Geauga_: Chardon Twp. (Wood, 1959:8).

_Greene_: Huffman Dam. _Hamilton_: Harrison; mouth of Miami River.

_Hardin_: "near" Hepburn. _Henry_: Maumee River, east of Napoleon;

Maumee River "near" Texas; Maumee River, 3 mi. W Texas. _Highland_:

Little Brush Creek, 2 mi. N Sinking Spring. _Huron_: Huron River

"near" Monroeville. _Jackson_: Canter's Cove, Jackson Twp.; Jackson

Lake. _Knox_: Brinkhaven. _Lake_: east branch Chagrin River, Kirtland;

Grand River, 4 mi. E Painesville. _Lawrence_: Pine Creek, Elizabeth

Twp. _Logan_: Miami River, "near" Indian Lake. _Lorain_: Oberlin.

_Lucas_: Lake Erie at Reno Beach, Jerusalem Twp.; Lake Erie, 1/2 mi.

offshore from mouth of Crane Creek; Maumee River at Maumee; Swan

Creek, W of Toledo; "near" Waterville; Swan Creek "near" Whitehorse.

_Madison_: London. _Medina_: Hinckley Lake. _Meigs_: Shade River,

below Darwin. _Miami_: Miami River, above Troy. _Monroe_: Cranenest

Fork, Green Twp. _Montgomery_: Mad River, Dayton; Miami River, Dayton;

Stillwater River, Dayton. _Morrow_: Kokosing River, Franklin Twp.

_Noble_: Jct. Sharon Twp. 1 and St. Rt. 78. (Adler and Dennis,

1960:27). _Ottawa_: East Harbour, Catawba Island. _Pike_: Chenoweth

Fork, Sunfish Twp.; Scioto River, Camp Creek Twp. _Ross_: Paint Creek

near Bainbridge. _Vinton_: Lake Hope; Lake Alma. _Warren_: Fort

Ancient. _Washington_: Dam No. 2, Muskingum River, "near" Marietta.

_Williams_: 1 mi. S Blakesley; St. Joseph River "near" Blakesley; West

Branch, St. Joseph River, Sec. 8, Bridgewater Twp.; Edgerton. _Wood_:

Grand Rapids; Grassy Creek, Rossford; Haskins; Maumee River opposite

Toledo.

PENNSYLVANIA: _Allegheny_: Monongahela River above McKeesport

(Atkinson, 1901:154); Ohio River at Pittsburgh (Wied-Neuwied _in_

Stejneger, 1944:44, 49). _Armstrong_: (Swanson, 1952:165). _Clarion_:

Clarion River "near" Clarion (Allen, 1955:228); Foxburg (= Foxbury?,

Boulenger, 1889:260). _Crawford_: _Elk_: _Erie_: Edinboro Lake.

_Forest_: (Swanson, _loc. cit._). _Indiana_: Plum Creek; Crooked Creek

(Netting _in_ Stejneger, 1944:48). _McKean_: (Swanson, _loc. cit._).

_Somerset_: Stoyestown (Surface, 1908:122). _Warren_: _Venango_:

Allegheny River south of Franklin (Swanson, _loc. cit._).

TENNESSEE: _Chester_: South Fork, Forked Deer River just E Henderson

(Endsley, 1954:40). _Clay_: Mill Creek, 3 mi. from Butler's Landing;

Obey River above mouth of Wolf River at Lilydale; mouth of Wolf River

(Shoup, Peyton and Gentry, 1941:75); Iron Creek "near" Willow Grove

(Stejneger, 1944:56). _Fentress_: _Jackson_: (Gentry, 1941:332).

_Lake_: Reelfoot Lake (Parker, 1948:29). _Obion_: Walnut Log (Parker,

1937:85); east shore of Reelfoot Lake, Samburg (Rhoads, 1895:386).

_Overton_: Medlock Branch, tributary of West Fork Obey River north of

Allred (Shoup, Peyton and Gentry, _loc. cit._). _Roane_: 2 mi. S

Kingston (Stejneger, 1944:55).

VERMONT: _Chittenden_: Lake Champlain, mouth of Winooski River; "near"

Burlington; Milton (= Minton) (Babcock, 1919:420). _Franklin_: Swanton

(Stejneger, 1944:55).

WEST VIRGINIA: _Randolph_: Tygart River at Elkins (Green, 1937:116).

WISCONSIN: _Burnett_: _Crawford_: (Pope and Dickinson, 1928:83).

_Dane_: Lake Wingra, Madison (Noland, 1951:54). _Grant_: (Pope and

Dickinson, _loc. cit._). _Green Lake_: Berlin (AMNH 6840-41, listed in

card file March 2, 1959). _Jefferson_: Lake Mills (Dickinson,

1950:75). _La Crosse_: West Salem (Pope, 1930:281). _Oneida_: _Pepin_:

(Pope and Dickinson, _loc. cit._). _Racine_: Eagle Lake (Edgren,

1944:498); Burlington; Rochester (Stille and Edgren, 1948:201).

_Sheboygan_: Sheboygan (KKA). _Trempealeau_: _Vernon_: "near" Viroqua

(Pope, _loc. cit._). _Walworth_: Lake Beulah (Dickinson, _loc. cit._).

_Washburn_: (Pope and Dickinson, _loc. cit._). _Waukesha_: Lac La

Belle (Cahn, 1929:8). _Winnebago_: Wolfe River (Dickinson, _loc.

cit._).

=Trionyx spinifer hartwegi= (Conant and Goin)

Western Spiny Softshell

Plates 35 and 36

_Amyda spinifera hartwegi_ Conant and Goin, Occas. Papers Mus.

Zool. Univ. Mich., No. 510:1, pl. 1, map 1, June 15, 1948.

_T[rionyx] s[pinifer] hartwegi_ Schwartz, Charleston Mus. Leaflet,

No. 26:11, May, 1956.

_Type._--Holotype, UMMZ 95365; alcoholic adult male; obtained at

Wichita, Sedgwick County, Kansas, in May, 1945, by Robert Young.

_Range._--Central United States in tributaries flowing into the

Mississippi River from the west, except the Red River drainage;

eastern Montana, North Dakota, and southern Minnesota south to eastern

Colorado, northern Oklahoma and Arkansas (see map, Fig. 19).

_Diagnosis._--Juvenal pattern of small ocelli, rarely as large as two

millimeters in diameter, or usually solid black dots that are not

much larger in center of carapace than at sides (mean OD/PL, Kansas,

.022); only one dark marginal line separating pale rim of carapace

from dorsal ground color.

_Description._--Plastral length of smallest hatchling, 2.8 centimeters

(USNM 9928); of largest male, 13.1 centimeters (USNM 55687); of

largest female, 25.5 centimeters (KU 2283).

Carapace olive, having small ocelli or black spots that are not much

larger in the center of the carapace than at the sides; pale rim of

carapace separated from darker ground color by one dark marginal line

and not four or five times wider posteriorly than laterally; large

females often having black dots at sides of carapace on mottled and

blotched pattern; pattern on snout of pale, dark-bordered stripes that

unite forming acute angle in front of eyes; well-defined dark markings

in subocular and postlabial region; pattern contrasting with ground

color on side of head; postlabial stripe broken, interrupted; pale

postocular stripe having blackish borders interrupted, not joining

with postlabial stripes; dorsal surface of soft parts of body having

contrasting pattern, largest blackish marks on hind limbs; elongate

tail of males having pale dorsolateral bands with well-defined, lower,

blackish borders; patterns on soft parts of body usually obscured or

absent on large females; underparts whitish, often having blackish

marks, except in center of plastral area; dark marks on webbing of

limbs, palms and soles; dark streaks often coincident with digits;

tubercles along anterior edge of carapace small and conical on adult

males, and conical or knoblike on large females; accessory, knoblike

tubercles in nuchal region and in middle of carapace posteriorly on

large females.

Ontogenetic variation in PL/HW, mean PL/HW of specimens having

plastral lengths 7.0 centimeters or less, 4.24, and exceeding 7.0

centimeters, 5.33; ontogenetic variation in CL/CW, mean CL/CW of

specimens having plastral lengths 8.5 centimeters or less, 1.12, and

exceeding 8.5 centimeters, 1.19; mean CL/PCW, 2.00; mean SL/HW, 1.30

(including subspecies _spinifer_); mean CL/PL, 1.38.

_Variation._--Variants include: CNHM 8949, UMMZ 72511 and TU 14591

having ocelli approximately 4 millimeters in diameter that are almost

solid spots; KU 17728 having pale stripes on snout that lack black,

inner borders; TTC 719 (female, plastral length 20.7 cm.), having

distinct pattern on snout; USNM 14535, 17823, 55684, and 123446 (from

different localities) having markings confined to margins of carapace

(Stejneger, 1944:66, suggested that USNM 17823 probably came from

Texas); UMMZ 92667 (female, plastral length 6.7 cm.) lacking pattern

on carapace.

_Comparisons._--_T. s. hartwegi_ can be distinguished from all other

subspecies of _T. spinifer_ by the presence of small dots and ocelli

on the carapace that are all of approximately the same size in

combination with only one dark marginal line. _T. s. hartwegi_

resembles _asper_ in having small blackish ocelli or dots on the

carapace but differs from _asper_ in having only one dark marginal

line. _T. s. hartwegi_ differs from _spinifer_ only in the small size

of the ocelli. _T. s. hartwegi_ resembles _spinifer_ and _asper_, but

differs from _pallidus_, _guadalupensis_ and _emoryi_ in having

blackish spots and ocelli on the carapace and lacking small whitish

spots. _T. s. hartwegi_ resembles _spinifer_, _asper_ and _pallidus_

but differs from _guadalupensis_ and _emoryi_ in having conical or

knoblike tubercles on the anterior edge of the carapace on large

females.

_T. s. hartwegi_ differs from the subspecies _asper_, _guadalupensis_

and _emoryi_ in having a narrower head, and from _emoryi_ in having a

wider carapace. _T. s. hartwegi_ resembles _spinifer_ and _asper_ but

differs from the other subspecies in having the carapace widest at a

plane approximately one-half way back on the carapace. _T. s.

hartwegi_ and _spinifer_ have longer snouts than do _pallidus_ and

_guadalupensis_ or _emoryi_. _T. s. hartwegi_ differs from _asper_ but

resembles the other subspecies in having a relatively longer plastron.

_Remarks._--The validity of _T. s. hartwegi_ has never been

questioned. It intergrades with _spinifer_ over a broad area

paralleling the Mississippi River. For convenience, specimens

occurring west of the Mississippi River are referred to the subspecies

_hartwegi_. Figure 8 shows much variation in size of ocelli on

different individuals from the same state. For example, UMMZ 92667,

plastral length 6.7 centimeters has a uniform pale brown carapace

lacking any dark marks, whereas UMMZ 92652, plastral length 5.9

centimeters has some ocelli three millimeters in diameter on the

carapace. Both are from Iowa. One specimen from Kansas, KU 1954

(Doniphan County, plastral length 11.8 cm.), has ocelli four

millimeters in diameter, and USNM 7648 captured farther west at Fort

Laramie, Wyoming, an adult male having a plastral length of 11.0

centimeters, has some ocelli five millimeters in diameter on the

carapace. TTC 1090, an adult male from the panhandle of Texas has some

ocelli so much as 5.5 millimeters in diameter. The size of the ocelli

seemingly varies in the same local population.

Specimens of _T. spinifer_ in the lower Mississippi Valley are

intergrades. Most individuals have small black dots on the carapace;

some have small ocelli (TU 7216, 7501, 11912, 12123-24) interspersed

with black dots (TU 5863), others have black dots confined to the edge

of the carapace (TU 157, 4539, 7105), and still others have no pattern

on the carapace (TU 7506, 13698.1, 10087.6). Two large males (TU

11580, 13025) have large ocelli (approximately five mm. in diameter)

that have nearly black centers. In general, there is more dark

pigmentation than farther north; some specimens have extensive

pigmentation on the ventral surface of the carapace and soft parts of

the body (TU 156, 5648). The dorsal surface of the limbs, especially

the hind limbs, have a bold, black marbling and may be almost

completely black (TU 5484, 5597). Many females, not exceeding plastral

lengths of 7.0 centimeters, have a pale blotched pattern of lichenlike

figures or have ill-defined black dots on the carapace (TU 10087,

13698.13, 13753.15).

Localities of specimens of _T. spinifer_ occurring in the Mississippi

River drainage in Mississippi are arbitrarily listed under the account

of the subspecies _spinifer_, whereas those in Louisiana (excluding

_pallidus_) are listed under the account of _hartwegi_.

Neither Over (1943) nor Wheeler (1947:169) record _T. s. hartwegi_,

respectively, from South Dakota or North Dakota; records from the

Missouri River drainage in Montana suggest the occurrence of the

species in that drainage in North and South Dakota.

_Specimens examined._--Total, 392 as follows: ARKANSAS: _Clay_: UMMZ

70735 (2), 7 mi. S St. Francis. _Crawford_: USNM 95352, Lee Creek, 7

mi. NW Natural Dam. _Drew_: CNHM 40785. _Lafayette_: KU 2225-29, 2944

(one of three specimens bearing last catalog number), 2963 (one of

three specimens bearing this catalog number), 2964 (one of two

specimens bearing this catalog number), Lewisville (see remarks under

the account of the subspecies _pallidus_). _Lawrence_: CNHM 8949;

CNHM 12598-600, 12602-04, TU 5855, UI 2413, Imboden; UI 2412, Black

River at Powhatan. _Marion_: TU 14591 (6), White River at Cotter.

_Prairie_: KU 1867, 1869, 1879, 1949-51, 2280-83, 2285-91 (2 specimens

bear catalog number 2287), 2307, 2761-62, 2666, 2826, 2842, 3346-47,

White River at DeValls Bluff. _Pulaski_: UMMZ 96540, Little Rock.

_Saline_: USNM 17823, Saline River at Benton. _Searcy_: UMMZ 92755,

Little Red River, 1.5 mi. SE Leslie. _Yell_: TU 14565, Petit Jean

Creek, 10 mi. N Casa. _County unknown_: CNHM 28566-67, Ouachita River.

IOWA: _Allamakee_: UMMZ 72556-58, 92642-49, Mississippi River "near"

Lansing. _Appanoose_: UMMZ 92667, Chariton River, 4.3 mi. N.

Centerville. _Decatur_: UMMZ 92651, Grand River, 3.5 mi. WSW Decatur.

_Dickinson_: UMMZ 55249, Milford; UMMZ 92655, Spirit Lake Twp.

_Hamilton_: USNM 9928, Webster City. _Hardin_: UMMZ 92650, Eldora.

_Louisa_: UMMZ 92654, Muscatine Slough, 12 mi. SW Muscatine, Muscatine

County. _Muscatine_: INHS 7675, 5.5 mi. SE Muscatine; USNM 54730-32,

Fairport. _Scott_: CNHM 433, Davenport; UMMZ 92656, Steamboat Slough,

2 mi. N Princeton. _Story_: UMMZ 92653, Squaw Creek at Ames.

_Washington_: UMMZ 92652, English River, 2 mi. E Riverside.

KANSAS: _Anderson_: KU 52286-87, 3-1/4 mi. E, 1/2 mi. N Colony.

_Atchison_: UMMZ 66939-41, Atchison. _Barber_: KU 17728, 4.5 mi. S Sun

City; KU 41379, 41742, 6 mi. N, 3.5 mi. E Sharon; USNM 100580,

Medicine River, 1 mi. S Lake City. _Cherokee_: KU 1323, Galena.

_Comanche_: KU 18385, 3-4 mi. SE Arrington. _Cowley_: UMMZ 75963, USNM

90441-44, 91022, 100529-30, "near" Winfield. _Doniphan_: KU 1943,

1952-54, Doniphan Lake. _Douglas_: KU 1955-56, Wakarusa River; KU

40176-77, Kansas River at Lawrence. _Franklin_: KU 3290. _Hamilton_:

KU 2990, Syracuse. _Harper_: KU 18159, 1 mi. N Harper. _Kingman_: USNM

95261, 2 mi. E Calista. _Labette_: KU 3339. _Lane_: KU 3738-41,

Pendennis. _Logan_: KU 16531, Smoky Hill River, 3 mi. SW Elkader.

_Meade_: KU 40210, Crooked Creek, 12.5 mi. S, 1-1/4 mi. W Meade.

_Montgomery_: KU 3731-32, Independence; KU 50856, Cherryvale Lake.

_Neosho_: UMMZ 69294, Caneville Creek, 32 mi. N. Parsons, Labette

County. _Osage_: KU 3294-96, Appanoose Creek. _Pratt_: KU 15931-32,

15934, State Fish Hatchery "near" Pratt. _Riley_: KU 48239, McDowell

Creek, WSW Manhattan; UMMZ 64434, "near" Manhattan. _Russell_: KU

3289. _Sedgwick_: UMMZ 95363-65, Wichita. _Shawnee_: USNM 123446,

Kansas River at Topeka. _Stafford_: KU 3758, Little Salt Marsh; KU

41743, 13.5 mi. N, 6 mi. E Stafford. _Trego_: KU 2757, 3769, Smoky

Hill River, 10 mi. N (NNE) Utica, Ness County; KU 51517, Saline River,

5 mi. N, 1/2 mi. E Wakeeney. _Wilson_: KU 56744-45, Verdigris River, 1

mi. S Altoona. _Woodson_: KU 55295, Neosho River, 1/2 mi. E, 1-1/2 mi.

S Neosho Falls. _County unknown_: USNM 51529.

LOUISIANA: _Catahoula_: TU 12629, Ouachita River, 4 mi. N

Harrisonburg. _Claiborne_: TU 13080, Caney Lake "near" Summerfield.

_Concordia_: KU 50849, Tensas River at Clayton; TU 16524 (3), USNM

012349, Lake Concordia; USNM 99865, Red River "near" Shaw. _East

Carrol_: TU 827-30, 905, 5644-45, Lake Providence. _Grant_: TU 12735,

Big Creek at Fishville, "near" Pollock. _Jefferson_: TU 5592-98, 7184,

10741, 10171, Mahogany Pond. _Lafourche_: TU 7105, 7132, 7216, 7501,

7505-07, 10087 (14), 11828-29, 11912, 11983 (2), 12123-28, 13502,

13679 (8), 13753 (22), 13766.2, Bayou Lafourche at Raceland.

_Morehouse_: USNM 11631 (2), Mer Rouge. _Natchitoches_: USNM 100420,

Cane River "near" Natchitoches. _Orleans_: TU 16169 (3), Audubon Park,

New Orleans; USNM 029310, "near" New Orleans. _Ouachita_: TU 12916,

12954, 12970-71, 13019, 13025, Bartholomew Bayou at Sterlington; TU

5988, Monroe. _Pointe Coupee_: TU 153, 156-59, 165, 5484, 5513,

5518-19, 5646, 5648, 5651, USNM 100202-12, False River at New Roads.

_Rapides_: TU 14040, Red River at Rapides. _Richland_: OU 25082. _St.

Bernard_: TU 16170, Delacroix Island. _St. Charles_: TU 4539, 4579,

5224, 5990, 11928 (12), 13698 (16), Bayou Gauche between Paradis and

Des Allemands; TU 5863, 11580, Bonnet Carre Spillway at Norco.

_Tensas_: TU 5762, Lake St. Joseph near Newellton. _Union_: USNM

138946, Meridian Creek, 1 mi. E Conway; USNM 138947, Ouachita River,

Alabama Landing. _Parish unknown_: MCZ 1622, Lake St. John (Concordia

or Tensas Parish); USNM 029266, Louisiana?

MINNESOTA: _Hennepin_: AMNH 4759-60, Fort Snelling. _Lesueur_: KU

46742-43, Waterville, Lake Tetonka. _Winona_: USNM 59263-66, Homer.

MISSOURI: _Carter_: UMMZ 70737, "near" Van Buren. _Chariton_: UI

17509, Triplett. _Franklin_: USNM 55689. _Gasconade_: UMMZ 95900,

Bourbeuse Creek, 8 mi. S Owensville. _Jefferson_: USNM 95405, Glaize

Creek. _Lewis_: USNM 59279-80, Canton. _Miller_: UMMZ 91929, Barren

Fork Tavern Creek, 5 mi. NW Iowna. _Newton_: UMMZ 82822, Shoal Creek,

12 mi. W Momit. _Phelps_: UMMZ 91930, Bourbeuse River, 10 mi. N St.

James. _Reynolds_: CNHM 35392, Black River at Warner Bay Spring; USNM

55688. _Ripley_: UMMZ 90435. _Shannon_: INHS 6223, Alley Spring State

Park. _St. Charles_: USNM 93089-94, Dardenne Creek, St. Peters. _St.

Louis_: USNM 55685-87, Mississippi River at St. Louis. _Stone_: USNM

55684. _Washington_: USNM 55690. _Wayne_: UI 16554, Sam A. Baker State

Park; UMMZ 95879, St. Francis River at Lodi. _County unknown_ (Wayne

or Butler): UMMZ 83264, Clark National Forest, St. Francis River.

MONTANA: _Big Horn_: USNM 54421, Crow Agency. _Roosevelt_: USNM 58,

Fort Union (locality reads "Yellowstone, Fort Union"; probably the

Yellowstone River near Fort Union). _Wheatland_: UMMZ 92005,

Musselshell River near Shawmut. _Yellowstone_: USNM 14535, Custer.

OKLAHOMA: _Alfalfa_: OU 9316, 2 mi. S Cherokee. _Cleveland_: OU 22973,

Norman. _Delaware_: UMMZ 81476, Spavinaw. _LeFlore_: OU 16802, 1.5 mi.

E Zoe. _Osage_: UMMZ 89628, Big Hominy Creek. _Pottawatomie_: OU

25175, 5 mi. SW Shawnee. _Rogers_: OU 7317, Verdigris River, 5 mi. W

Claremore; UMMZ 81473-74, near Garnett, Tulsa County; UMMZ 81475, 4

mi. NE Inola. _Sequoyah_: OU 9008, 2 mi. NE Gore; TU 13885, Little

Vian Creek, 1 mi. E Vian. _Texas_: OU 5005, 5 mi. SE Guymon. _Tulsa_:

TU 17061, Bird Creek "near" Skiatook, Osage County. _Woods_: CHNM

11809, Waynoka; OU 9432, 2.5 mi. W Waynoka; OU 9579, 9581-82, 1 mi. S

Waynoka.

TEXAS: _Hansford_: TTC 719, 10 mi. S, 2 mi. W Gruver. _Hutchinson_:

TTC 1090, Carson Creek, Turkey Track Ranch.

WYOMING: _Goshen_: USNM 7648, Fort Laramie. _Weston_: UMMZ 78080,

Beaver Creek.

NO DATA: CNHM 21687-88, 22925. SM 142 (locality of Waco, McLennan

County, Texas, believed in error). USNM 7649, 11625, 19622-23, 36412

(Illinois River).

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