ACCOUNTS OF SUBSPECIES

=Geomys bursarius lutescens=, Merriam

_Geomys bursarius lutescens_ Merriam, North Amer. Fauna, 4:51,

October 8, 1890; Scheffer, Technical Bull., U. S. Dept. Agric.,

224:6, January, 1931.

_Geomys lutescens_ Merriam, North Amer. Fauna, 8:127-29,

January 31, 1895; Lantz, Trans. Kansas Acad. Sci., 19:175,

1905; Lantz, Kansas State Agric. College Bull., 129:335, April,

1905; Hibbard, Trans. Kansas Acad. Sci., 36:240, 1933; Black,

30th Bienn. Rept. Kansas State Board Agric., 35:182, 1937;

Swenk, Missouri Valley Fauna, 2:1, February 1, 1940; Allen,

Kansas State Teachers College, Emporia, Bull. Inf. in Educ., 20

(no. 5):15, May, 1940; Hooper, Occas. Papers Mus. Zooel., Univ.

Michigan, 420:3, June 28, 1940.

_Geomys lutescens lutescens_, Hibbard, Trans. Kansas Acad.

Sci., 47:74, 1944.

_Type locality._--Sandhills on Birdwood Creek, Lincoln County,

western Nebraska.

_Distribution in Kansas._--Northwestern Kansas, eastward

certainly to Ellis County, southward certainly to Scott County.

_Description._--Animals with total length averaging no more

than 272 mm.; length of vertebrae of tail averaging no more

than 92; hind foot averaging no more than 35. Color: In autumn

pelage, upper parts Light Ochraceous-Buff becoming Buckthorn

Brown in middorsal region and there forming a faint

longitudinal band; sides Pale Yellow Orange. In summer,

Buckthorn Brown on upper parts with a dorsal band, especially

distinct on specimens from Ellis and Trego counties; specimens

from farther west lack the distinct dorsal band. Underparts

Gray Drab and sometimes whitish, usually whitish in young

specimens; basal color of pelage Deep Neutral Gray; fore and

hind feet whitish. Skull: Zygomatic arch broadly and squarely

spreading anteriorly; temporal impressions uniting to form a

low sagittal crest in adult males, but in adult females and in

young males the impressions usually remain apart; shape of

interparietal varying from subquadrate in young specimens to

subtriangular or triangular in adults; in some young specimens

the interparietal is reduced to a minute, ovoid bone.

_Comparisons._--See comparisons in the accounts of other subspecies

occurring in Kansas.

_Remarks._--In his monographic revision of the pocket gophers, Merriam

(1895:129) recorded 3 "typical or nearly typical" specimens from Trego

County, and 18 "non typical" specimens as follows: Garden Plain,

Sedgwick County, 4; Belle Plain, Sumner County, 5; Cairo, Pratt County,

6; Kiowa, Barber County, 2; and Ellis, Ellis County, 1. A detailed

discussion of Merriam's account of the distribution of _Geomys

lutescens_ in Kansas is given by Swenk (1940:11-12).

Judging by specimens in the University of Kansas Museum of Natural

History, _G. bursarius lutescens_ in Kansas is restricted to the

northwestern part of the state, reaching southward certainly to Scott

County and eastward certainly to Ellis County; precise limits of

distribution of this subspecies are unknown. Additional collecting is

necessary to determine where the range of _lutescens_ meets the ranges

of the other subspecies. The specimens studied are remarkably uniform.

One specimen obtained in October, in Trego County, is slightly lighter

colored than any other from Kansas. In other characteristics it agrees

with specimens from northwestern Kansas and from the type locality.

_Specimens examined._--Total number 32, as follows: _Cheyenne

County_: 23 mi. (by road) NW St. Francis, 3. _Rawlins County_:

2 mi. NE Ludell, 10. _Logan County_: 5 mi. W Elkader, 3; no

locality more precise than county, 1. _Trego County_: Wakeeney,

4; 12 mi. S Collyer, Perrington Ranch, 3; no locality more

precise than county, 5. _Scott County_: 4 mi. S Scott City, 2.

_Ellis County_: Hays State College Campus, Hays, 1.

=Geomys bursarius majusculus= Swenk

_Geomys bursarius majusculus_ Swenk, Missouri Valley Fauna,

1:6, December 5, 1939; Hibbard, Trans. Kansas Acad. Sci.,

47:74, 1944.

_Geomys bursarius_, Baird, Expls. and surveys for a railroad

route from the Mississippi River to the Pacific Ocean, pt. 1,

Mammals, 377, 1857; Merriam, North Amer. Fauna, 8:120, January,

1895; Lantz, Trans. Kansas Acad. Sci., 19:175, 1905; Lantz,

Kansas State Agric. College Bull., 129:335, April, 1905;

Scheffer, Kansas State Agric. College Ento. and Zooel. Dept.

Bull., 172:199, September, 1910; Hibbard, Trans. Kansas Acad.

Sci., 36:240, 1933; Allen, Kansas State Teachers College

Emporia Bull. Inf. Stud. in Educ., 20 (no. 5):15, May, 1940.

_Geomys bursarius bursarius_, Black, 30th Bienn. Rept. Kansas

State Board Agric., 35:181, 1937.

_Geomys breviceps_, Baird, Expls. and surveys for a railroad

route from the Mississippi River to the Pacific Ocean, pt. 1,

Mammals, 380, 1857.

_Type locality._--Lincoln, Lancaster County, Nebraska.

_Distribution in Kansas._--Northeastern Kansas, westward

certainly to Clay and Marion counties and southward certainly

to Greenwood County.

_Description._--Color: Upper parts Mummy Brown in fresh

appearing pelage of February but in more worn pelage of March

more reddish being near (16') Prout's Brown; top of head and

sometimes back darker than rest of upper parts; underparts

usually with some whitish anteriorly; fore and hind feet and

approximately distal half of tail white. Size: Large, total

length averaging more than 280 mm. in males and 257 in females;

hind foot averaging 35 mm. or more in males. Skull: Large;

rostrum averaging more than twice as long as wide; sagittal

crest high in males and barely present in females; occiput

vertical when skull is laid top down; least width of braincase

less than

distance from alveolus of upper incisor to middle of lateral

border of P^4 at alveolar border.

_Comparisons._--From _Geomys bursarius lutescens_, _majusculus_ differs

as follows: Color darker, Mummy Brown to Prout's Brown instead of

Buckthorn Brown. In both sexes: head and body a fifth to a sixth longer;

hind foot 5 to 6 per cent longer; skull averaging larger in all parts

measured except that premaxillae (in each subspecies) extend equally far

posteriorly to nasals; diastema longer in relation to basilar length;

rostrum longer relative to its width; sagittal crest higher; rostrum

often more depressed distally; angle of suture between maxilla and jugal

more obtuse.

From _G. b. bursarius_, according to Swenk (1939:6), _majusculus_

differs in larger size.

From _G. b. illinoensis_, _majusculus_, according to Komarek and Spencer

(1931:405), differs in brownish instead of slate-gray coloration and in

two cranial characters as follows: Nasals straight-sided instead of

shaped like an hour-glass, and superficial canals on palatine extending

anteriorly beyond first molar, and from there anteriorly more or less

separated. The first of these characters does not always hold;

occasional individuals of _majusculus_, for example some from Douglas

County, have the nasals shaped like an hour-glass.

From _G. breviceps dutcheri_, _majusculus_ differs in larger size (hind

foot more than 33 mm. in males, and 29 in females; basilar length more

than 42 mm. in males and 36 in females); dorsal exposure of jugal longer

than width of rostrum measured between ventral margins of infraorbital

foramina.

From _G. bursarius major_ of southcentral Kansas (for example Harvey

County), _majusculus_ differs in slightly darker color, being Mummy

Brown instead of Prout's Brown; size larger (in males total length more

than 284 mm., hind foot 35 or more, basilar length of skull more than

42, and in females total length 265 or more, hind foot averaging 33 or

more, and basilar length 40 or more).

Skull: Averaging larger in all parts measured, except that premaxillae

do not extend so far posteriorly to nasals in either males or females;

interorbital constriction slightly narrower in adult females; temporal

ridges forming a more prominent sagittal crest in adult males (sagittal

crest barely present in some adult males of _major_ from Harper County).

_Remarks._--In employing the subspecific name _majusculus_ we are

following Swenk (1939:6) who on the basis of larger size differentiated

the animals from southeastern South Dakota, the eastern parts of

Nebraska and Kansas, and the western and southern parts of Iowa, from

_G. bursarius bursarius_ to which he assigned a more northern geographic

range. In the absence of comparative materials of the northern

subspecies we cannot make an independent decision on the validity of

_majusculus_ and recognize that if it is inseparable from _G. b.

bursarius_ the latter name will apply to specimens from northeastern

Kansas. We are the more uncertain about applying the name _majusculus_

to specimens from eastern Kansas because they average smaller than

topotypes. Only at the northeasternmost locality in Kansas (3 mi. N

Cummings, Atchison County) do specimens average as large as topotypes of

_majusculus_. Farther southward they become progressively smaller in

eastern Kansas, and we interpret this as intergradation with the still

smaller subspecies _major_, to the southwest. The average external

measurements of two adult males from Atchison County are: 321-99-35.

Thirty-six miles farther south, in Douglas County, 16 adult males

average 289-80-36. From Hamilton, Greenwood County, 80 miles farther

southwest, nine adult males average 284-83-35. The maximum total length

recorded at these three localities is: Atchison County, 342 (1 of 2

specimens), Douglas County, 308 (1 of 16 specimens), Greenwood County,

357 (in coll. of Dr. Glenn C. Rinker and 1 of 15 males of all ages

involved). It will be seen, therefore, that although there is a trend to

smaller average size toward the southward, the maximum of 357

millimeters total length at Hamilton exceeds the maximum of 352

millimeters recorded by Swenk (1939:3) among 86 males at Lincoln where

the recorded average is largest.

Four specimens from Salina (Debold Farm) are intermediate structurally,

as they are also geographically, between _G. b. majusculus_ on the one

hand and _Geomys bursarius lutescens_ and _Geomys bursarius major_ on

the other hand. In color they agree with _majusculus_, as they do also

in width of nasals posteriorly, in more obtuse angle of the rostrum and

maxillary arm of the zygomatic arch. They agree with _G. b. lutescens_

in having the occiput inclined anterodorsally, and are intermediate

between _majusculus_ and _lutescens_, but nearer the latter in size of

skull and in length of the rostrum relative to its width.

_Specimens examined._--Total number, 148, as follows: _Clay

County_: 6 mi. SW Clay Center, 3. _Jackson County_: 10-1/2 mi.

WSW Holton, 1; no locality more precise than county, 1.

_Atchison County_: 3 mi. N Cummings, 2. _Jefferson County_:

Oskaloosa, 1. _Leavenworth County_: Fort Leavenworth

(Government Hill, 2; Engineer Hill, 1), 6; no locality more

precise

than county, 19. _Saline County_: Salina, Debold Farm, 4 (coll.

of A. J. Kirn). _Morris County_: 1-1/2 mi. N Council Grove, 3.

_Douglas County_: 1 mi. NW Midland, 2; 1 mi. N Lawrence, 1;

2-1/2 mi. W Lawrence, 2; 1 mi. W K. U. Campus, 2; 1 mi. W

Lawrence, 2; 1/2 mi. W Lawrence, 2; "W K. U. Campus," 2; K. U.

Campus, 4; Lawrence, 23; South Lawrence, 1; 1/2 mi. SW K. U.

Campus, 2; Southwest K. U. Campus, 1; Haskell Institute, 1;

4-1/2 mi. S Lawrence, 1; 7 mi. SW Lawrence, 6; 7-1/2 mi. SW

Lawrence, 1; 8 mi. SW Lawrence, 1; 10 mi. S Lawrence, 1; 11 mi.

SW Lawrence, 3; no locality more precise than county, 15.

_Marion County_: 1-1/2 mi. NE Lincolnville, 6; 4 mi. SE

Lincolnville, 1; 6 mi. S Lincolnville, 1. _Greenwood County_:

Hamilton, 1; 1/2 mi. S Hamilton, 4; 1 mi. S Hamilton, 4; 4 mi.

S and 14 mi. W Hamilton, 6; 8 mi. SW Toronto, 1; 8-1/2 mi. SW

Toronto, 5; no locality more precise than county, 6.

=Geomys bursarius jugossicularis= Hooper

_Geomys lutescens jugossicularis_ Hooper, Occas. Papers Mus.

Zooel., Univ. Michigan, no. 420: 1, June 28, 1940; Hibbard,

Trans. Kansas Acad. Sci., vol. 47, p. 75, 1944.

_Type locality._--Lamar, Prowers County, Colorado.

_Distribution in Kansas._--Extreme southwestern part of state,

northward certainly to Hamilton County and south certainly to

Morton and Seward counties.

_Description._--A yellowish-cinnamon colored animal, with body

of medium size, zygomatic plate of maxilla deep and mastoid

process small.

_Comparisons._--Differs from _Geomys bursarius industrius_ in slightly

lighter color; occiput not strongly inclined anterodorsally.

From _G. b. lutescens_, _jugossicularis_ differs in less buffy

coloration and deeper zygomatic plate of maxilla.

_Remarks._--_G. bursarius jugossicularis_ and _G. bursarius industrius_

intergrade in the southern part of Meade County. Some specimens from

this area show a coloration resembling that of _G. b. jugossicularis_;

nevertheless, one specimen from Morton County has the occiput

anterodorsally inclined as in _G. b. industrius_.

Specimens examined from Hamilton County correspond closely to _G. b.

jugossicularis_; they agree with it both in color and in cranial

characters.

_Specimens examined._--Total number, 20, distributed as

follows: _Hamilton County_: 1 mi. E Coolidge, Conard Farm, 4.

_Morton County_: 12 mi. NE Elkhart, 2; Cimarron River, 12 mi. N

Elkhart, 4; no locality more precise than county, 6. _Seward

County_: 1 mi. E Arkalon, 4.

=Geomys bursarius industrius=, new subspecies

_Geomys lutescens_ Merriam, North Amer. Fauna, 8:127, January

31, 1895.

_Geomys breviceps llanensis_, Hibbard, Trans. Kansas Acad.

Sci., 36:240, 1933; Black, 30th Bienn. Rept. Kansas State Board

Agric., 35:181. 1937.

_Geomys lutescens jugossicularis_ Hooper, Occas. Papers Mus.

Zooel., Univ. Michigan, 420:1, June 28, 1940.

_Type._--Male, adult, skin and skull, no. 14083 Museum of

Natural History, University of Kansas; from 1-1/2 miles north

of Fowler, Meade County, Kansas; obtained December 30, 1941, by

H. H. Hildebrand, original number 16.

_Distribution in Kansas._--Southwestern Kansas from Meade

County eastward certainly to Pratt and Clark counties; from

Pawnee County southward probably to the Oklahoma boundary.

_Diagnosis._--Size of body medium; color of upper parts Cinnamon

Brown; skull with occiput strongly inclined anterodorsally in

males.

[Illustration: FIG. 2. Three views of the skull of the type

specimen of _Geomys bursarius industrius_. A. Lateral view; B.

Dorsal view; C. Ventral view. All natural size.]

_Description._--Color: Upper parts Cinnamon Brown, slightly

reddish, but in some specimens collected in September, in

Pawnee County, near (15' _i_) Ochraceous-Tawny; underparts

usually Wood Brown, somewhat whitish anteriorly; forefeet

white; hind feet and approximately distal half of tail whitish.

Size: Medium (see measurements), total length averaging not

more than 271 mm. in males and 254 in females; hind foot

averaging not more than 35 mm. in males and less than 32 in

females. Skull: In males, least width of braincase equal to

distance from alveolus of incisor to anterior border of

alveolus of first upper molar, occiput strongly inclined

anterodorsally, temporal impressions usually united in a low

sagittal crest, zygomatic arch heavy and curved at level of

jugal bone. In adult females least width of braincase

approximately equal to distance from alveolus of incisor to

anterior border of alveolus of first upper molar (not

premolar); occiput less inclined anterodorsally

than in males; temporal ridges not forming a sagittal crest. In

young females the width of the braincase is more than the

distance between the alveoli of the incisor and first molar.

_Comparisons._--_G. lutescens industrius_ differs from _G. lutescens

lutescens_ in: Color darker; least width of braincase not equal to

(usually more than) the distance from the alveolus of incisor to the

anterior border of the alveolus of the first upper molar.

_G. lutescens industrius_ differs from _G. lutescens jugossicularis_ in:

Color slightly darker, the former being Cinnamon Brown instead of

Vinaceous Cinnamon, with hairs basally Deep Neutral Gray in upper parts

and underparts. Skull: Jugular part of zygomatic arch more curved

(convex dorsally) and occiput far more inclined anterodorsally; lower

part of mastoidal ridge more prominent.

For comparison with _G. l. major_, see account of that subspecies.

_Remarks._--Judging from the known specimens of this subspecies, it has

the smallest geographic range of any of the subspecies in Kansas, but

additional collecting in Hodgeman County and counties to the north and

west of it may extend the known range in those directions; collecting in

Comanche County and in adjoining parts of Oklahoma may extend the known

range to the southward.

The anterodorsal inclination of the occiput in males is the one cranial

character in which _industrius_ differs from all of the subspecies with

adjoining geographic ranges. The existence of this unique (among

adjoining subspecies) cranial character is the principal reason for

according subspecific status to this animal. Although it has other

characters which are fairly uniform over a considerable geographic area,

these other characters, namely, Cinnamon Brown color of the upper parts

and medium size of the body, after all, are conditions intermediate

between those in _jugossicularis_ to the west and those in the darker

and larger animals assigned to _major_ to the eastward. Considering the

intermediate geographic position of _industrius_, the color and size are

approximately what a person would predict by study of only the animals

to the west and those to the east. Therefore, the color and size

probably are indicative of intergradation between _jugossicularis_ and

_major_. Still, there is the anterodorsally inclined occiput in

males--a character of a unique sort--and this influences us to give

subspecific status to this animal with full recognition of the fact that

it is a "weak" subspecies as compared with any one of the adjoining

subspecies.

Hooper (1940:2) in naming as new _Geomys lutescens jugossicularis_

referred to his new subspecies a skin-only from Meade County State Park.

Our more abundant material from there shows the cranial conformation to

be that of _industrius_ to which we accordingly assign the specimens.

However, with only a skin available, we, too, would have used the name

_jugossicularis_ because the color is paler than in other specimens of

_industrius_ and this paleness indicates intergradation between the two

named subspecies. Specimens from Pratt County are slightly darker than

_industrius_ thereby indicating intergradation between _industrius_ and

_major_.

_Specimens examined._--Total number, 58, distributed as

follows: _Pawnee County_: Jct. Pawnee and Arkansas rivers,

Larned, 6; 1 mi. S and 1 mi. E Larned, 7. _Edwards County_: 1

mi. W and 3-1/2 mi. S Kinsley, 1. _Kiowa County_: Rezeau Ranch,

5 mi. N Belvidere, 2. _Pratt County_: Pratt, 14; no locality

more precise than county, 1. _Meade County_: 3-1/2 mi. NE

Fowler, 2; 2 mi. N Fowler, 2; 1-1/2 mi. N Fowler, 2; 1-1/4 mi.

N and 3/4 mi. E Fowler, 2; 7 mi. N Meade, Cudahy Ash Pit, 2; 13

mi. SW Meade, 9; State Lake, 2; State Park, 4. _Clark County_:

7 mi. SW Kingsdown, E. A. Stephenson Ranch, 1; 6 mi. S

Kingsdown, 1.

=Geomys bursarius major= Davis

_Geomys lutescens major_ Davis, Texas Agric. Exp. St., Bull.

no. 590:32, August, 1940; Hibbard, Trans. Kansas Acad. Sci.,

47:75, 1944.

_Geomys lutescens_ Merriam, N. Amer. Fauna, 8:129, January 31,

1895.

_Geomys breviceps llanensis_, Lantz, Trans. Kansas Acad. Sci., 20

(pt. 2): 215, 1907; Hibbard, Trans. Kansas Acad. Sci., 36:240,

1933; Black, 30th Bienn. Rept. Kansas State Board Agric.,

35:182, 1937; Swenk, Missouri Valley Fauna, 2:12, February 1,

1940.

_Type locality._--Eight miles west of Clarendon, Donley County,

Texas.

_Distribution in Kansas._--Southcentral Kansas, northward

certainly to Ellsworth County, westward certainly to Stafford

and Barber counties and eastward to Cowley County.

_Description._--Color: Upper parts varying from Brussels Brown

in some specimens to nearly Prout's Brown, especially in

specimens from central part of state. Top of head, and

sometimes back, darker than rest of upper parts, but no well

defined black stripe; underparts varying from whitish to nearly

Buffy Brown; fore and hind feet and approximately distal half

of tail white. Size: Large (see measurements). Skull: Sagittal

crest absent in females and barely present in males; least

width of braincase more than distance from alveolus of incisor

to middle of lateral border of P^4 at alveolar border. Length

of auditory bulla (from anteroventral edge of paroccipital

process of exoccipital to hamulus of peterygoid), in each sex,

more than 8 mm.; occiput usually vertical when skull is laid

top down; zygomatic arch broadly and squarely spreading,

divergent anteriorly; rostrum averaging less than twice as long

as wide.

_Comparisons._--From _G. bursarius lutescens_, _major_ differs in color

darker, premaxillae extending slightly farther posteriorly; temporal

impressions usually forming a more well-marked sagittal crest in males;

ventral side of zygomatic arch, at level of jugal bone, more curved.

From _G. bursarius majusculus_, _major_ differs in slightly lighter

color, smaller size of body; in males, total length less than 284 mm.;

hind foot 34 or less; basilar length of skull less than 42; in females

total length less than 264, hind foot no more than 33, and basilar

length less than 39.

From _G. bursarius industrius_, _major_ differs in color, being Prout's

Brown, instead of Cinnamon Brown (less Fuscous); body averaging 10 per

cent longer; total length in males from 9 to 9.7 per cent longer, hind

foot 9.7 per cent longer on the average; skull averaging larger in all

parts measured. Occiput less inclined anterodorsally; top nearly flat,

less arched than that of _G. b. industrius_; auditory bulla averaging

slightly larger and less inflated.

_Remarks._--Specimens of this subspecies from Norman, Cleveland County,

Oklahoma, and Canton, Dewey County, Oklahoma, and most of those from

Kansas, are more Fuscous than topotypes and tend toward _G. bursarius

majusculus_. Specimens from McPherson County have a darker dorsal stripe

resembling that of _G. bursarius majusculus_. One adult from Little Salt

Marsh, Stafford County, is pale, closely resembling topotypes.

Most of the cranial characters, nevertheless, are constant in all

available specimens, except that in specimens of each sex from the type

locality the basilar length averages 4 to 5 per cent shorter. In the

constancy of size of the relatively large auditory bullae and in the

nearly flat dorsal profile of the cranial part of the skull, the

specimens from Kansas agree with the specimens from the type locality.

Specimens from Harper County have the occiput slightly inclined

anterodorsally and thus are reminiscent of _industrius_ which has an

even greater inclination of the occiput. Probably the appearance in

dilute fashion of this character in Harper County is properly to be

interpreted as intergradation with _industrius_. If so, the actual

intergradation may be to the northwest _via_ Pratt County since

specimens from Barber County, immediately west of Harper and lying

between Harper County and the range of _industrius_, do not have the

occiput so inclined.

Of a pair of adults from eight miles west of Rosalia, Butler County,

the female is indistinguishable in color from adults of _G. b.

industrius_ from northern Meade County and from two specimens from

eleven miles west of Clarendon, Donley County, Texas, near the type

locality of _G. b. major_. The male from eight miles west of Rosalia is

darker as compared either with _G. b. industrius_ or _G. b. major_ and

the coloration of the upper parts resembles those in _G. b._

_majusculus_; the underparts are only slightly paler than the upper

parts as in _majusculus_. Measurements of the skulls are intermediate

between the averages for _G. b. majusculus_ and those for _G. b. major_.

These specimens from eight miles west of Rosalia are intermediate

structurally, and since they are intermediate geographically between _G.

b. majusculus_ and _G. b. major_, they suggest intergradation of the two

subspecies. The specimens in question are referred to _major_ because

the size is nearer that of _major_. It is mainly the intermediate nature

of these two specimens from Butler County, and the intermediate nature

of the specimens from McPherson County, Kansas, that have caused us to

treat _G. b. majusculus_ as only subspecifically distinct from the more

western subspecies, _major_.

_Specimens examined._--Total number, 77, as follows: _Ellsworth

County_: 2 mi. S Ellsworth, 1. _McPherson County_: Smoky Hill

River, 1 mi. S and 1/2 mi. W Lindsborg, 5; 1/2 mi. E McPherson,

1. _Stafford County_: Little Salt Marsh, 12; no locality more

precise than county, 3. _Reno County_: 8 mi. N and 1 mi. E

Haven, 2. _Harvey County_: 1 mi. E and 1/2 mi. N Halstead, 1;

Halstead, 3. _Butler County_: 8 mi. W Rosalia, 2. _Barber

County_: near South Bridge, Sun City, 1; 2 mi. S Sun City, 1;

Wells Ranch, Aetna, 5; "1 mi. W Aetna," 3; near South Bridge,

Aetna, 1; near Bridge, 1 mi. S Aetna, 2. _Harper County_: 4-1/2

mi. NE Danville, 8; 1 mi. N Harper, 11; 3 mi. S Harper, 1.

_Cowley County_: 3 mi. SW Arkansas City, 4; 3 mi. SE Arkansas

City, 9; 3 mi. S Arkansas City, 1.

Subspeciation in Pocket Gophers of Kansas, [KU. Vol. 1 No. 11] · 第 7 / 11 题