HISTORY

The first published reference that we have found to pocket gophers of

Kansas is Prof. Spencer F. Baird's (1857:377, 380) mention of two

specimens from Fort Riley. One he identified as _Geomys bursarius_ (p.

377) and the other (p. 380) he doubtfully referred to _Geomys

breviceps_. Both specimens were obtained by Dr. W. A. Hammond. J. A.

Allen (1874:49) reported pocket gophers from Kansas under the generic

name "Geomys?". Professor M. V. B. Knox (1875:21) published a list of

Kansas mammals in which he used the names _Geomys bursarius_ Shaw and

_Geomys breviceps_ Baird, the last one for the specimen taken by Dr.

Hammond, at Fort Riley. Baker (1889:57) employed the name _Geomys

bursarius_ Rich. for the gopher "found along the hundredth meridian,

between N latitude 38 deg. 30' and 39 deg. 30'." He reported this animal as

common in western Kansas. Merriam (1895:129) recorded _G. bursarius_ and

_G. lutescens_ from Kansas. Allen (1895:265) recorded five specimens of

_Geomys lutescens_ collected between September 16 and October 13 at Long

Island, Phillips County, Kansas, by W. W. Granger. Since that time

several papers, some of them dealing mostly with habits of pocket

gophers, have been published in which reference is made to _Geomys_ in

Kansas. Hibbard (1933:240) recognized three species: _G. bursarius_, _G.

lutescens_, and _G. breviceps llanensis_. In 1944 (74-75) he recorded

_Cratogeomys_ from Meade County, on the basis of two skulls dug out of

the ground, and he recognized the same three full species of the genus

_Geomys_ that he did in 1933, along with two additional subspecies.

Specimens to the total number of 335 from Kansas have been available

for the present study of the five subspecies recognized. The reason for

arranging all of the named kinds as subspecies of a single species is

that intergradation has been found to occur between every pair of kinds

having contiguous geographic ranges. The characters previously thought

by some writers constantly to differentiate, say, _Geomys lutescens_ of

western Kansas from _Geomys bursarius_ of eastern Kansas, prove not to

do so; instead, in areas geographically intermediate between the

geographic ranges of the two kinds, the pocket gophers are intermediate

in morphological characters and therefore are regarded as intergrades.

Intergradation of this kind here is accepted as the criterion of

subspecies, and lack of such intergradation as the criterion of species.

Search for structural characters, distinctive of the different kinds,

additional to those characters noted by other writers, has resulted in

the finding of a few such characters but they too are subject to

intergradation. Therefore the several kinds are arranged as subspecies

of a single species which takes the name _Geomys bursarius_ because it

is the oldest available name. Detailed comment on specimens showing

intergradation are to be found in the accounts of _G. b. bursarius_ and

_G. b. major_.

Subspeciation in Pocket Gophers of Kansas, [KU. Vol. 1 No. 11] · 第 5 / 11 题