HELOTES OCTOLINEATUS. _Jen._

_H. corpore lineis longitudinalibus nigricantibus octo; pinnis

dorsali, anali, caudalique, maculis fuscis; vertice striis elevatis

duabus subparallelis; preoperculo distinctè denticulato, et ad

marginem limbi internum subcristato; operculo mucronibus duobus,

superiore minimo; squamis ubique lævissimis._

B. 6; D. 12/9; A. 3/7; C. 17, &c.; P. 15; V. 1/5.

LONG. unc. 9. lin. 9.

FORM.—Body oblong. Greatest depth exactly four and a half times in the

entire length. Length of the head rather less than the depth of the

body. Snout short and obtuse. Jaws exactly equal: each with a

broadish band of velutine teeth, which are all, apparently even the

most minute, three-pointed, although this character is not very

obvious except in the outer row, which are longer than the others.

No vomerine teeth appear externally, but they may be felt through

the skin of the palate, and on dissecting this off, there is brought

to view a small hard disk rough with minute asperities. Mouth very

little cleft, the commissure not extending more than half-way

between the end of the snout and the anterior margin of the orbit.

Eyes rather large; their diameter one-fourth the length of the head.

Maxillary, when the mouth is closed, concealed in part beneath the

suborbital, the lower margin of which is somewhat sinuous and

obscurely denticulated, the denticulations being concealed by the

membrane and more easily felt than seen. The denticulations on the

preopercle very manifest. The principal spine on the opercle slender

and very sharp, not exactly straight, but slightly curved, the

convexity of the bend being downwards; above is a second spine, but

very small and easily overlooked. The crown of the head has two

nearly parallel elevated lines, which take their origin between the

nostrils, and terminate at the occiput, but do not meet as in the

_H. sexlineatus_;[6] a third line commences there exactly between

them, and runs singly in a backward direction down the middle of the

nape; this last is scarcely more than half the length of the two

former. The cheeks and pieces of the gill-cover are scaly; but not

the cranium, snout, jaws, or limb of the preopercle, which last is

margined internally by a slightly elevated ridge. The scales on the

body are thin and small, and without any trace of denticulations on

their free edges, even under a magnifier, and the body of the fish

is quite smooth to the touch rubbed either way. Lateral line as in

_H. sexlineatus_. Dorsal also nearly similar, but more deeply

notched, the membrane beyond the eleventh spine falling nearly to

the base of the twelfth, which precedes the soft portion: sixth

spine longest, equalling very nearly, but not quite, half the depth;

the eleventh equals the second; the twelfth is about one-third

longer than the eleventh, but is itself scarcely half the first soft

ray. The anal has three soft rays less than the _H. sexlineatus_,

and there are apparently but two spines, the first being (at least

in this specimen, where, however, there may have been a portion

broken off) quite short and rudimentary; the second and third spines

are both slender, the former being rather more than half the length

of the latter, and this last rather more than half the first soft

ray. All the fins take their origin as in _H. sexlineatus_. The

pectorals are about two-thirds the length of the head. The ventrals,

which are very near together, are longer than the pectorals, but do

not equal the head: they have no elongated scale between them, or in

their axillæ.

COLOUR.—For the most part similar to that of the _H. sexlineatus_; but

the longitudinal dark lines are more numerous, amounting to eight,

with faint traces of a ninth: the additional ones are on the upper

half of the sides, or above the lateral line, there being four

(instead of two) above that one which passes through the eyes; the

sixth extends the whole length of the fish from the end of the

maxillary to the base of the caudal; the seventh passes immediately

below the pectoral, and terminates in advance of it, without quite

reaching to the edge of the gill-cover; the eighth is exactly

equidistant from the pectoral and ventral; this last is a very

narrow pale line, but the others, with the exception of the first

two, are broader and well marked. The soft portion of the dorsal, as

well as the anal and caudal are spotted; the spots on this last

unite to form transverse fasciæ; those on the anal are not very

well-defined. The pectorals and ventrals are without spots, and

pale.

Habitat, S. W. coast of Australia.

This species was procured in King George’s Sound, New Holland. It

closely approaches the _H. sexlineatus_ of Cuvier and Valenciennes, the

only species of the genus hitherto described, and obtained in the same

seas by MM. Quoy and Gaimard. I have little hesitation, however, in

pronouncing it to be distinct. Independently of the additional

longitudinal lines on the body, and the spots on the fins, which, it is

expressly stated by the above authors, are not present in the _H.

sexlineatus_, it is distinguished by the striæ on the crown not meeting

behind, the dorsal being rather more deeply notched, and the fin-ray

formula different. There is one soft ray less in the dorsal, and three

less in the anal; also the first spine in this last fin, if it be not

broken off in this specimen, is quite rudimentary. The scales present no

trace of cilia on their free edges.

1. PINGUIPES FASCIATUS. _Jen._

PLATE V.

_P. corpore fasciis transversis duodecim castaneo-fuscis, alternis

latioribus; dentibus palatinis paucis, conicis, subaggregatis, vix

seriem formantibus; pharyngalibus aculeiformibus; membranâ

branchiali mediocriter emarginatá; operculo spinâ unicâ forti,

alterâ superiore obsoletâ; pinnis ventralibus accuratè thoracicis._

B. 6; D. 7/27; A. 1/24; C. 15, &c.; P. 18; V. 1/5.

LONG. unc. 12. lin. 9.

FORM.—Body thickest, as well as deepest, in the region of the

pectorals, compressed behind, and becoming more so as it approaches

the tail; depth also gradually diminishing from that point. The

greatest depth is rather less than five and a half times in the

entire length: head contained four times and a quarter in the same.

The thickness at the pectorals is at least three-fourths of the

depth: and the thickness of the head is quite equal to it. Dorsal

line nearly straight from behind the eyes, in front of which the

profile descends obliquely. Eyes high, nearly reaching to the line

of the profile; a little behind the middle point of the length of

the head; their diameter rather less than one-fifth of this last;

the distance between them one diameter and a half. The commissure of

the lips does not reach to the eyes by a space equalling half the

diameter of the eye. Jaws equal. Lips very thick and fleshy, and

partially reflexed, like those of a _Labrus_. Teeth very similar to

those of that genus. In the upper jaw, an outer row (extending all

round) of strong, sharp, slightly curved teeth, regularly set, and

nearly even, but with the anterior ones a little the longest; in all

about forty, twenty on each side; behind these a velutine band,

broadest in front, but also extending the whole way round. In the

lower jaw, a row of curved strong teeth, similar to those above, but

extending only half-way along the sides of the jaws (about nine on

each side), and followed by about seven or eight short blunt conical

ones; a broad velutine band behind the longer curved teeth, but not

behind the others. On the front of the vomer are four or five large

blunt conical teeth, mixed with smaller ones of the same form: there

is also a small group of these little conical teeth at the

commencement of each palatine, but they are not carried on further

in a single row.[7] Tongue small and inconspicuous, fastened down

except just at the tip, smooth. Pharyngeal teeth in strong card; but

no conical ones behind that are visible. Branchial membrane united

to its fellow, and free all round at the margin, with a moderately

deep notch underneath. Preopercle rounded at the angle; the

ascending margin oblique. Opercle with a strong sharp spine at its

upper angle, but not extending beyond the membrane; a second

rudimentary one above it obtusely rounded. Small scales on the

cheeks, preopercle, and opercle, but not on the snout, or between

the eyes, or on the suborbital, or jaws, or branchial membrane, or

interopercle. The scales on the body are rather small, finely

ciliated on their edges, thin, and of an oblong form, cut square at

the basal margin, with a fan of twelve or fifteen striæ. Lateral

line not very strongly marked, taking nearly a straight course from

the upper part of the scapular to the caudal. No particular lines,

markings, or pores, about the head, jaws, or between the eyes.

Pectorals rounded; two-thirds the length of the head. Ventrals

exactly beneath them, a very little shorter, thick and fleshy, so

that the rays can hardly be distinguished. Dorsal and anal similar

to those of the _P. Brasilianus_; the former has the spinous rays at

first low, but the rest of the fin is of one uniform height,

equalling a little less than half the depth: the latter commences

under the sixth soft ray of the dorsal, and terminates in the same

line. Caudal square, with rows of small scales between the rays for

half their length: also a few minute scales at the base of the

pectoral rays, but none on the other fins.

[Illustration:

_Fish. Pl. 5._

_W. Hawkins del^t._

_Pinguipes fasciatus. ¾ Nat. Size._

]

COLOUR.—“Above pale ‘chestnut brown,’ so arranged as to form

transverse bands on the sides; sides, head, fins, with a black

tinge; beneath irregularly white: under lip pink: eyes with pupil

black, and iris yellow.”—D. _In spirits_; the back and upper half of

the sides are brown, the lower half of the sides and belly pale,

with twelve transverse dark fasciæ, the alternate ones broader than

the others. The dorsal and anal appear to have been bluish, the tint

increasing in intensity from the base upwards; but the former is

edged above with a narrow white line just beneath the tips of the

rays, which extends the whole length of the soft portion of the fin.

The inside of the ventrals appears also to have been bluish; but the

pectorals are pale, or yellowish. Caudal brown like the back.

Habitat, coast of Northern Patagonia.

From the east coast of Patagonia, in Lat. 37° 26′. There can be no doubt

of its belonging to the genus _Pinguipes_, with which it agrees in its

very strong resemblance to the _Labridæ_, as regards the head, lips, and

teeth, and in its fleshy ventrals; but there are very few teeth on the

palatines, seeming to show that there is not much ground for separating

this genus from _Percis_. In many of its characters, it resembles the

_P. Brasilianus_ of Cuvier, but it is decidedly distinct in others. It

differs slightly in its proportions; in the palatine and pharyngeal

teeth; in the position of the ventrals, which are not at all jugular,

but immediately beneath the pectorals; in the branchial membrane being

more notched; and in having two soft rays less in the anal. The colours

also are different.

This fish is so like a _Labrus_, that at first sight it might easily

deceive a student. Nevertheless its vomerine teeth, spines on the

opercle, and ciliated scales, point out its right family. At the same

time no system can be considered natural, which does not admit

_Pinguipes_ as one of the connecting links between the _Percidæ_ and

_Labridæ_.

The zoology of the voyage of H.M.S. Beagle, [vol. 4 of 5] · 第 44 / 282 题