_Enemies._--Mr. Quillin writes to me: “While this species is still

fairly abundant in southern Texas, it was much more plentiful ten or

more years ago. Because of the damage the birds wrought to telephone

and telegraph poles, the various concerns owning such property secured

passage of a law placing all woodpeckers on the unprotected list.

This done, they gave section crews of the railroads shotguns, and the

killing was on in earnest. Hunters and others helped, and the result

has been a marked decrease in the ranks of this species. The killing,

or controlling still continues. However, pressure is now being brought

to place the birds back on the protected list, and this will be done

sooner or later. There is no getting around the fact that the birds did

cause considerable damage. In this species we have a woodpecker which

for centuries had been pecking into hard mesquite trees. Along came the

soft pine poles and these same birds immediately literally ate them

up. I have seen 16 holes, three of which were deep enough for nesting

sites, in one small pole, not over 10 inches in diameter.”

DISTRIBUTION

_Range._--North-central Texas south to Central Mexico; nonmigratory.

The golden-fronted woodpecker ranges =north= to central Texas (San

Angelo and Dallas). =East= to Texas (Dallas, Giddings, Cuero, Corpus

Christi, and Brownsville); Tamaulipas (Matamoros, San Fernando, Ciudad

Victoria, and Tampico); southeastern San Luis Potosi (Valles); Hidalgo

(Ixmiquilipam and Tula); and the Federal District of Mexico (near

Mexico City). =South= to the Federal District of Mexico (near Mexico

City); Michoacan (Querendero, Morelia, and Patzcuaro); and Jalisco

(Ocotlan and Guadalajara). =West= to Jalisco (Guadalajara); Zacatecas

(Calvillo, Aguas Calientes, and Chicalote); northwestern Durango

(Boquilla, Sestin, and Rosario); eastern Chihuahua (Julimes); and

central Texas (Eagle Pass, Fort Clark, Kerrville, and San Angelo).

_Egg dates._--Texas: 66 records, March 30 to June 29; 33 records,

April 24 to May 17, indicating the height of the season.

CENTURUS UROPYGIALIS UROPYGIALIS Baird

GILA WOODPECKER

PLATES 32-34

HABITS

In the desert regions of our southwestern borders, this gay little

woodpecker is one of the commonest, noisiest, and most conspicuous

birds, always much in evidence, and always seeming to protest, in

whining tones, the intrusion of strangers. Its center of abundance

seems to be on the great desert mesas of southern Arizona, where the

infertile soil is scantily covered with a scattered growth of creosote

bushes, low mesquites, an occasional cholla or barrel cactus and

dotted with single specimens or little groups of the giant cactus, or

saguaro. But it is also common in the river bottoms, covered with a

heavier growth of mesquite, and in the canyons of the foothills among

the cottonwoods, willows, and sycamores. It ranges from an elevation of

2,500 feet on the mesas up to 4,000, or even 4,500, feet in the canyons

and foothills.

In this region, it is a dominant species and a very useful neighbor,

even if unintentionally, for the many species of birds and small

mammals for which it provides homes. M. French Gilman (1915) puts it

very well as follows:

Were it not for the Gila woodpecker (_Centurus

uropygialis_) what would become of the several species

of birds that use already prepared cavities for their

domiciles? In some cases these tenants do not even

await the pleasure of the excavators, but take forcible

possession. In holes excavated by Gila woodpeckers there

may regularly be found nesting the elf owl, ferruginous

pigmy owl, ash-throated flycatcher, and Arizona crested

flycatcher. Occasionally a cactus wren makes use of the

handy hollow, and once I saw one occupied by a Lucy

warbler. A big “rough-neck” scaly lizard frequents the

holes when not too high in the cactus, and in two holes in

willow trees I found snakes. It is not pleasant to insert

one’s hand and have a big lizard or snake crawl up the

arm to escape. Rats and mice are sometimes found in the

deserted holes, especially if the tree be much decayed

and with cracks and hollows connecting holes at different

heights in the tree or branch. So these woodpeckers may be

considered among the class of innocent or unintentional

benefactors.

In addition to the species mentioned by Mr. Gilman above, we found

saguaro screech owls, desert sparrow hawks, and western martins

nesting in the old holes made by woodpeckers. Some of these holes were

doubtless made by Mearns’s gilded flickers, perhaps those that were

used by the larger species, as this woodpecker is fairly common in

the same region and nests regularly in the saguaros. These old holes

make ideal nesting sites, for the sap of the cactus hardens around the

excavations, making them fairly permanent nesting boxes; I have seen

these gourd-shaped pockets still persisting in fallen saguaros, where

the pulp had all rotted away, leaving only the skeleton ribs of the

dead giant.

_Nesting._--While collecting with the late Frank C. Willard in

southern Arizona in 1922, we examined seven occupied nests of the Gila

woodpecker. The first of these was found on May 17, at Fairbank, in the

valley of the San Pedro River; the nest was a cavity 15 inches deep

in a dead branch of a cottonwood, 15 feet above ground. Five days,

May 19 to 23, were spent in Pima County, in the vicinity of Tucson,

between the mesquite forest in the valley of the Santa Cruz River and

the southern end of the Santa Catalina Mountains. Two nests were found

in the mesquite forest on May 19, both in mesquite trees, one 20 and

one 25 feet from the ground; one contained only a single fresh egg and

the other held a brood of young. We had an interesting experience here

the next day. While crossing the forest, I saw a Gila woodpecker fly

out from what I supposed was its nesting hole, about 15 feet up in a

mesquite stub; the bird made such a great fuss about it that I felt

sure that we had a set of woodpecker’s eggs within easy reach, and I

called Mr. Willard to investigate it. He climbed the stub and chopped

out the hole, while the woodpecker was flying about, scolding us and

showing the greatest concern. But, much to our surprise, he pulled out

an elf owl and three unmistakable elf owl’s eggs. I killed the owl

and shot the woodpecker, which still seemed much interested; and, on

skinning and sexing both specimens, I found that the woodpecker was a

male and the owl a female. We were naturally much puzzled to figure out

the relationship between the two birds and their interest in the nest.

But, since reading Mr. Gilman’s remarks, quoted above, that sometimes

the woodpecker’s tenants “do not even await the pleasure of the

excavators, but take forcible possession,” it has occurred to me that

probably this was a case in point. The owl may have appropriated the

finished burrow of the woodpecker, and the latter was trying to evict

an unwelcome tenant.

The remaining four nests found in this vicinity, and one found by

Mr. Willard on June 11, were all in saguaros on the desert mesa; the

heights from the ground varied from 16 to 20 feet; and the cavities

varied in depths from 15 to 20 inches; there was one set of five eggs,

two nests held four and one three eggs; and in one nest were two young

and an addled egg.

Referring to the nesting habits of this woodpecker in the vicinity of

the Gila River, in Arizona, Mr. Gilman (1915) writes:

Nesting sites in this locality are restricted to giant

cactus (_Cereus giganteus_), cottonwood and willow, as they

are the only suitable material for a nest excavation. More

nests are found in the giant cactus, as these plants are

more numerous than the others, and more “peckable,” though

the willows and cottonwoods along the river and the canals

are well patronized when sufficiently decayed. Of the nests

I examined I should say that fifty per cent were in the

cactus, and the rest equally divided between the other

trees mentioned. * * *

As to the size of the holes in the cactus as compared with

those in cottonwood and willow, I found no appreciable

difference. I expected the holes in the cactus to average

a little larger owing to possible greater ease in

excavating but the difference was too slight to be sure

of in measuring. Of eighteen holes measured, the average

diameter was 1.95 inches; the largest was 2.25 inches and

the smallest 1.87 inches. The deepest hole was 16 inches,

with the entrance 2 inches in diameter. The shallowest one

was 9 inches, with entrance a little less than 2 inches in

diameter. The average depth of holes measured was a little

more than 12 inches. Many of the holes were not exactly

circular, there being a difference of from ⅛ to nearly ½

inch between the long and short diameter if it be allowable

to use the term in that way. Usually the nest hole runs

straight in for a short distance before turning downward,

the distance seemingly depending on the texture of the

wood. In one case the hole went straight back for nine

inches before turning downward. It was in a big cottonwood

stump, and the bird excavated horizontally until decayed

wood was reached, when the hole turned downward. This was

an extreme case, as the depth horizontally is usually about

three inches. In the giant cactus it varies according to

the diameter of the trunk, the smaller the trunk the less

distance before turning downward. * * *

The same nest hole is used more than one season, both in

cactus and other locations. In 1913 I found a nest in a

big cottonwood stump containing young. The next year it

had young again, and I cut into it to measure the hole and

count them.

Frank C. Willard (1912) says: “I think it is their habit to dig fresh

holes after raising their brood of young. These fresh holes are not

occupied that year but are made use of the next year when the sap has

had a chance to dry and form the hard lining which coats the inside of

all the cavities. I have found but one fresh hole occupied as a nest.”

Bendire (1895) also says that “most of their nesting sites are used for

several years in succession; in fact, I doubt very much if a freshly

excavated hole in a giant cactus is fit to nest in the same season.

Both sexes assist in excavating the nesting site.”

In the heavily incrusted nest cavity in a giant cactus, the eggs lie

on the bare, hard floor of the nest, there being no chips to furnish a

soft bed.

In addition to the trees mentioned above, the Gila woodpecker has been

found nesting more rarely in oaks and palo-verdes.

_Eggs._--The Gila woodpecker lays three to five eggs, three or four

being much oftener found than five. The eggs are pure white and

not very glossy when fresh, but sometimes quite glossy when heavily

incubated; they vary from ovate to elliptical-ovate and are sometimes

quite pointed. The measurements of 52 eggs average 25.14 by 18.56

millimeters; the eggs showing the four extremes measure =27.43= by

18.80, 26.6 by =20.1=, =22.86= by 17.27, and 23.9 by =16.6= millimeters.

_Young._--Incubation is said to last about two weeks, and is probably

shared by both parents. Mr. Gilman (1915) writes:

It is not easy to determine just what food the young in

the nest are given, but insects play a prominent part, as

I have seen them frequently carried to the young. Fruit

is also used, as I watched one parent carry ripe Lycium

berries several times to the nest; after emerging from the

hole she would halt at the entrance each time and “lick her

chops.” * * *

The young are fed by the parents for a long time after

leaving the nest, and they are regular little beggars. One

pair stayed around our house for several months, and became

quite tame. They were missed during the breeding season

but soon came back with three youngsters to share the good

things found on the bird tables in the yard. The young,

though as large as their parents, would flutter their

wings and sit with open beak as though the old ones told

them to “open your mouth and shut your eyes,” etc. The old

ones would try to get them to eat watermelon placed on the

tables, but the babies would not be shown; the parents had

to put it in their mouths. They followed the parents from

perch to perch, begging for food until I expected to see

them chastised. The pair in question stayed with the three

juvenals until they had them broken to eat for themselves,

and then left. After a proper interval they came back with

two more young ones, thus indicating that a second brood is

sometimes raised. The abundant supply of food may have been

a determining factor in the number of broods raised.

_Plumages._--The nestlings are naked and blind at first but become

fully clothed in the juvenal plumage before leaving the nest. The young

male, in juvenal plumage, is much like the adult male, but the colors

are generally paler, the head and under parts grayer, the barring on

the upper parts less distinct, and the white bars are suffused with

brownish buff; the red patch on the crown is smaller and often consists

of only a few red feathers; and the bill is somewhat smaller and

weaker. The young female is like the young male but has no red on the

head. I have been unable to trace the postjuvenal molt, but young birds

in the following spring are apparently like the adults. Adults have a

complete postnuptial molt in August, September, and October.

_Food._--Major Bendire (1895) says: “Its food consists of insects of

various kinds, such as ants, beetles, grasshoppers, and larvæ, and in

season largely on the sweet, fig-like fruit of the sahuaras, the giant

cactus, and also, to a considerable extent, on the viscous berries of

a species of mistletoe which is commonly found on most of the larger

cottonwoods, oaks, and mesquite trees in these regions. These sticky,

whitish-looking berries are a favorite food of many Arizona birds.”

Mr. Gilman (1915) writes:

The food, of this woodpecker is varied, nearly everything

being grist that comes to his mill. He pecks around decayed

and dying trees as well as green ones, and presumably get

the insects usually found and eaten by such birds. The

giant cactus is pecked into very frequently, and I believe

some of the pulp is eaten. The small punctures made are not

enlarged, and in some cases quite an area is bitten into.

The fruit of the giant cactus is eaten as long as it lasts,

and the berries of the Lycium are also freely eaten. The

Gila woodpecker frequents corn fields, and pecks through

the husks into the ears of corn. The birds may peck in

at first to get a worm, but it is a case similar to the

discovery of roast pig as portrayed by Lamb. They alight on

the ground and feed upon table scraps thrown to chickens,

three of them being regular morning visitors, star

boarders, to a pen of chickens I fed. They are very fond of

peaches and pears, and volubly resent being driven from a

tree of the fruit. They peck holes in ripening pomegranates

and then the green fruit beetle helps finish the fruit They

relish grapes, both white and colored, and will spear one

with their bill and carry it to a convenient crevice where

it may be eaten at leisure. On bird tables I have tried

them with various articles of food and found very little

that they rejected. They would not eat cantaloupe at all

but were regular watermelon fiends, eating it three times

a day and calling for more. They did not care for oranges,

and I had no success in trying to teach them to eat ripe

pickled olives. I tried the olive diet on them because two

Mocking-birds in our yard learned to eat this fruit. Meat,

raw and cooked, was eaten, and they ate suet greedily.

Their favorite cut of beef was the T-bone steak and we

always left some meat on the bone for them. They picked it

clean, and if a new supply was slow in coming the softer

parts of the bone were devoured. * * * Mr. Frank Pinkley,

custodian of the Casa Grande Ruins told me of a pair of

these woodpeckers that stayed around his home and became

quite tame, coming into the shed to drink from a can of

water. He said they got into the habit of sucking the eggs

in the chicken house, or at least pecking into them and

eating of the contents. * * *

The Indians store corn in the ear on the flat tops of their

houses and sheds, * * * and each home has one or more of

woodpecker retainers or pensioners hanging about most of

the time. This corn provides an abundant and sure source

of food, and the birds make the most of it. I have never

seen any indication of food-storage on the part of the Gila

woodpecker, as with the California Woodpecker, for they

live in a claw-to-beak fashion. They peck at a kernel until

it comes off the cob, when it is carried to a post or tree

and placed firmly in a crack. Here it is pecked to pieces

and eaten. They seem never to swallow a kernel whole but

always break it up.

W. L. Dawson (1923) says that this woodpecker indulges in “a systematic

search for birds’ eggs, especially those of the Lucy warbler, yellow

warbler, and Arizona Least Vireo. In case of the first-named, the

eggs are devoured in spite of the most emphatic protests of the tiny

parents; but eggs of Cardinal, Cooper Tanager and Towhee must be

obtained by stealth.”

A. H. Anderson (1934) writes:

In the Tucson, Arizona, area a gall-insect (_Pachypsylla

venusta_) frequently attacks the leaves of the hackberry

tree (_Celtis reticulata_). The galls form on the leaf

petiole, becoming from a quarter to half an inch in

diameter. During the winter the outer shell hardens like a

nut.

I have often seen the Gila woodpeckers tear the galls loose

from the twigs and, flying to a fence post, proceed to

chisel out the contents. The hard gall is wedged into a

crack on the post and then opened by repeated hammering.

Around the base of one fence post I counted nearly 300

empty shells. Sometimes cracks in nearby trees are used. At

one time five of these woodpeckers were seen in a single

tree, all of them feeding on the galls.

_Behavior._--The Gila woodpecker is not only the most abundant

woodpecker, in fact one of the most abundant birds, in the region it

inhabits, but it is more conspicuous, noisier, and more active than

any of its neighbors. It is always much in evidence, always protesting

the intrusion of a stranger, and shows the greatest concern when

its nest is approached, especially if it has young. It is a close

sitter and will often remain in the nest hole to peck viciously at an

investigating hand; while the nest is being robbed, it flits nervously

about, scolding vociferously with all the vile epithets it can muster.

As to its behavior with other species, Mr. Gilman (1915) writes:

This woodpecker has not the best disposition in the world,

for he is very quarrelsome and intolerant. He fights his

own kin and all the neighbors that he dares. He, or she,

is a great bluffer however and when “called”, frequently

side-steps, subsides, or backs out entirely. I saw one

approach a Bendire Thrasher that was eating, and suddenly

pounce on him. He had the thrasher down and I was thinking

of offering my friendly services as a board of arbitration,

when the under bird crawled from beneath and soon gave

the woodpecker the thrashing of his career. Several times

I have seen the woodpeckers start to attack Bendire and

Palmer thrashers, but they were always bluffed or beaten at

the game. With the Bronzed Cowbirds it is a drawn battle,

sometimes one and then the other backing down. Most other

birds, such as Cardinals, Abert Towhees, Dwarf Cowbirds and

Cactus Wrens do not attempt to assert their rights, but

always take a rear seat. But when it is woodpecker versus

woodpecker it seems not to be a case of “Thrice armed is he

who hath his quarrel just”, but rather, “Four times he who

gets his blow in fust”.

I had two bird tables about twenty feet apart, and

frequently one woodpecker might be peacefully assimilating

watermelon, when another one would come hurrying up and

make a dive at him, causing a retreat to the other table.

Frequently the new-comer would then follow and drive him

from the second table. He seemingly would rather fight than

eat if another was eating at the same time. One day I saw

him, or her, I forget which, hanging to the edge of the

table busily eating steak, when another one perched on the

table and made a vicious stab at him. He dodged backward

clear under the table, though retaining his hold, and then

bobbed up again, just like the Punch and Judy show. The

attack was renewed, and the dodging as well, but this time

he did not “come back”. Another day one of them was at

work on a piece of melon when one of his fellows came and

perched on the end of the table. The diner made a pass at

the new comer, and seizing him by the feathers of the neck

held him suspended over the end of the table for a few

seconds.

_Voice._--Major Bendire (1895) says: “Its ordinary call note, sounding

like ‘dchürr, dchürr,’ can be heard in all directions in the spring;

when flying from one point to another it usually utters a sharp, shrill

‘huit’ two or three times, resembling the common call note of the

Phainopepla, and which may readily be mistaken for it. It is also more

or less addicted to drumming on the dead tops of cottonwood, sycamore,

and mesquite trees.”

Mr. Gilman (1915) writes:

As a neighbor, the Gila Woodpecker is permanently on the

map, and is afraid neither of being seen nor heard. He is

much in the public ear with a variety of notes and calls.

His sociable conversational notes somewhat resemble those

of the California Woodpecker but are shriller. In such of

his notes as are directed at humanity there is a peevish

complaining tone, especially if closely approached when

feeding on fruit or some other delicacy. In such cases

there is only one term that exactly describes his attitude

and utterances, and that is the phrase “belly-aching.”

In fact all of his talk at us has a distinctly “colicky”

tone and one feels like giving him something to whine

about. His ordinary call slightly resembles that of the

Flicker but is not quite so loud; altogether he is quite a

conversationalist.

_Field marks._--The Gila woodpecker should be easily recognized as a

medium-sized woodpecker, about the size of a hairy woodpecker, with a

grayish-brown head, neck, and under parts and a back narrowly barred

with black and white; in flight a white patch shows in the wing and

basal half of the primaries, and the black and white barring on the

central tail feathers is rather conspicuous; the red crown patch of the

male is conspicuous only at short range.

_Fall._--This woodpecker is apparently somewhat given to wandering in

fall and spring, for W. E. D. Scott (1886) says that he does not see it

about his house, at an elevation of 4,500 feet in Pinal County, Ariz.,

in summer, but that it is rather common there in fall and spring.

DISTRIBUTION

_Range._--Southwestern United States and western Mexico; nonmigratory.

The range of the Gila woodpecker extends =north= to extreme southern

Nevada (Clark County); southern Arizona (Sacaton, Rock Canyon, and

Tombstone); and southwestern New Mexico (Red Rock and probably Gila).

=East= to New Mexico (probably Gila); eastern Sonora (Fronteras, Boca

de Huachy, and Nuri); southwestern Chihuahua (Batopilas); western

Durango (Chacala); and western Zacatecas (Calvillo). =South= to

southwestern Zacatecas (Calvillo); and Jalisco (Guadalajara, Santa

Ano, and Rio Ameca). =West= to Jalisco (Rio Ameca); Nayarit (Tepic and

San Blas); southwestern Sinaloa (Escuinapa, Labrados, and Mazatlan);

Baja California (Cape San Lucas, Santa Margarita Island, San Ignacio,

Rosario, San Quintin, Las Palmas, and the Alamo River); southeastern

California (Calexico, probably Brawley, Palo Verde, and Needles); and

southern Nevada (Clark County).

This species has been separated into three geographic races, or

subspecies. Typical _C. u. uropygialis_ is the form found in that part

of the range lying in the United States, and this race also is the one

found in the western mainland of Mexico. The cardon woodpecker (_C. u.

cardonensis_) is found in the northern part of Baja California south

to about latitude 28° N. Brewster’s woodpecker (_C. u. brewsteri_)

occupies the cape district of Baja California north to San Ignacio and

including also Santa Margarita Island.

_Egg dates._--Arizona: 26 records, April 7 to May 30; 13 records,

May 5 to 25, indicating the height of the season.

Baja California: 10 records, April 21 to June 2.

CENTURUS UROPYGIALIS CARDONENSIS Grinnell

CARDON WOODPECKER

HABITS

In describing and naming this race, Dr. Joseph Grinnell (1927a) says:

In its main characters similar to _Centurus uropygialis

uropygialis_, but general coloration much darker: whole

head (except for red patch on crown) and anterior lower

surface strongly tinged with snuff brown rather than pale

drab; and white barring on closed wings, tail, dorsum,

rump, flanks, and lower tail coverts, narrower, leaving

the black-barring correspondingly broader. Similar to _C.

u. brewsteri_, but size larger, and coloration darker, in

the same respects though not to quite so great a degree as

shown in comparison with _uropygialis_. In other words, the

new form differs from both the previously known races in

the deeper brown tinge of the head and lower surface and in

the greater degree of predominance of black over white in

the barring.

Life histories of North American woodpeckers · 第 31 / 43 段