They do not show a preference for any one kind of tree but

are found on both living and dead shrub oaks, long-leafed

pines, loblolly pine, sycamore, sour gum and sweet gum.

They work on both trunks and limbs but usually at low

heights, from the ground up to twenty feet above. On a

vertical surface these birds work up, spiraling it and

tapping it as they go. They move by a series of short

hops, propping themselves each time with their tails. When

hopping lightly along a horizontal limb they still use

their tails as props. Perhaps their most astonishing feat

is to spiral horizontal limbs, and to cling beneath them

and hammer them with their backs down. Sometimes they work

their way up to the very tip of slender shoots.

Even in a heavy wind they cling to the violently swaying

twigs while eating, but they stay only a short time before

flying to a tree trunk to perch and rest before trying it

again. * * *

The flight of these birds is strong and undulating, with

fast beating wings, and generally only from one tree to

the next. Where the trees are not very close together,

they swoop down to within a few feet of the ground and

then fly with nearly level flight until they glide up to

their next stopping place. Where they have to fly out

across intervening open fields their flight becomes more

undulatory, at times deeply so.

DRYOBATES VILLOSUS HARRISI (Audubon)

HARRIS’S WOODPECKER

PLATES 4, 5

HABITS

The range of this well-marked subspecies is now restricted to the humid

coast belt from southern British Columbia southward to Humboldt County,

Calif. In 1895, Bendire wrote:

Until within the last few years all the Hairy Woodpeckers

from the eastern slopes of the Rocky Mountains to the

Pacific coast have been considered as belonging to this

subspecies. * * *

The breeding range of this race, as now considered, is a

very limited one, and is probably coextensive with its

geographical distribution. It is apparently confined to

the immediate vicinity of the coast, and is not found at

any great distance inland. Among the specimens collected

by me at Fort Klamath, Oregon (mostly winter birds),

there are two which might be called intermediates between

this and the more recently separated _Dryobates villosus

hyloscopus_, but the majority are clearly referable to

the latter. In the typical Harris’s woodpecker the under

parts are much darker, a smoky brown, in fact; it is also

somewhat larger and very readily distinguishable from

the much lighter-colored and somewhat smaller Cabanis’s

woodpecker.

Dawson and Bowles (1909) say: “Doctor Cooper judged the Harris to be

the most abundant Woodpecker in Western Washington; and this, with

the possible exception of the Flicker (_Colaptes cafer saturatior_),

is still true. The bird ventures well out upon the eastern slopes of

the Cascade Mountains, and is found sparingly in the higher mountain

valleys; but its favorite resorts are burns and the edges of clearings,

rather than the depths of the woods.”

Johnson A. Neff (1928) quotes Dr. I. N. Gabrielson as saying: “The

Harris woodpecker is found throughout western Oregon from the western

slope of the Cascades to the coast, altho in the Rogue River Valley

some specimens which are close to ‘orius’ have been taken. I have one

labeled ‘orius’ by Dr. H. C. Oberholser, also have typical Harris from

this district, so that this is probably the region of intergradation

between these two forms.”

Harris’s woodpecker, like many other races of the humid Northwest coast

region, is darkly colored, one of the most easily recognized of all the

hairy woodpeckers. Even in Audubon’s day it was recognized, described,

figured, and named by him in honor of his friend Edward Harris. Ridgway

(1914) describes it as “similar to _D. v. hyloscopus_, but under parts

(including lateral rectrices) light drab or buffy drab-gray, instead of

white or nearly white, the head-stripes and stripe on back also usually

more or less suffused with the same color, often uniformly light drab;

average size slightly larger.”

_Courtship._--Theed Pearse (1934) gives us the following interesting

description of the courtship display of this woodpecker:

There were two males on the limb of a small cedar and my

attention was first drawn to them by their note, which is

very similar to the flicker’s “wickety” note but softer,

and might almost be described as “caressingly soft.” Both

birds were calling.

The displaying bird would draw in its head so that no neck

was apparent, with beak pointed outwards and upwards and

would then slowly swing the upper part of the body from

side to side, thereby bringing into play the red nape

marking. Once the bird very rapidly lifted its wings into

an upright position, at other times there was a quivering

flicking motion of the wings as they lay against the sides.

The two birds flew to another branch, settling side by

side and instantly each froze, the neck drawn in and the

beak pointed upwards. They were perched sideways on the

branch and were displaying the white markings in the tail

which each of them would slightly spread and turn out

(to one side). The feathers of the back were at the same

time hunched up as though to show up the white markings

there also. They did this several times before flying off

together.

_Nesting._--Although this is evidently a common bird within its

restricted range, surprisingly little has appeared in print regarding

its nesting habits. Authentic eggs seem to be very rare in collections;

most of the eggs in collections, of which I have the records, that are

labeled _harrisi_, prove to be referable to one of the neighboring

subspecies.

D. E. Brown writes to me:

Its nesting cavities may be at any height from 4 feet to

well over 100 feet from the ground. I found a nest 8 feet

up in an 8-inch dead fir stub in a dry open locality. The

female flushed from the nest, and the date was just right

for fresh eggs, May 6. The cavity was carefully opened. It

was 16 inches deep and contained a single egg. This egg

was so fresh and the shell so clear that the yolk could be

plainly seen. The cavity was carefully repaired with bark

from the stub, held in place with black thread. Both birds

were near all this time, complaining loudly. I returned in

five days. The bark was still in place, but the egg was

gone. The dust from the decaying stub, where the egg had

been, was very dry, and I am of the opinion that the parent

bird had removed the egg.

G. D. Sprot has sent me a beautiful photograph (pl. 4) of a nesting

site of this woodpecker in a dead alder stump in a coniferous forest

clearing, near Mill Bay, Vancouver Island, taken May 23, 1928.

Dawson and Bowles (1909) say:

The nest of this bird is usually placed well up in a small

dead fir tree in some burn or slashing on dry ground.

It is about ten inches deep and has no lining save fine

chips, among which the crystal white eggs, four or five in

number, lie partially imbedded. Incubation is begun from

the last week in April to the last week in May, according

to altitude, and but one brood is raised in a season.

These Woodpeckers are exceptionally valiant in defense of

their young, the male in particular becoming almost beside

himself with rage at the appearance of an enemy near the

home nest.

S. F. Rathbun sends me the following note on a Harris’s woodpecker that

made an attempt to dig a nesting hole in a small young fir topped about

10 feet up: “The tree had been cut off so that it could be used as one

of the supports of a cross piece to which a swing was attached. The

woodpecker began to dig a hole in the topped upright, and the owner of

the place called me up and wanted to know what the bird was and what

it was up to. I told him all about it and suggested that he keep away

from the stub. Two weeks later, I asked him how the bird was getting

on. He said at first the bird was busy digging away every day, but of

late seemed to have something the matter with it, for ‘lately every

day it just sat with its head sticking out of the hole and did no work

on it.’ I cut a piece from the edge of the entrance and quickly found

out. When the crossbar for the swing had been nailed to the sapling,

a 10-inch spike was used to hold it; and this had gone nearly through

the sapling. The woodpecker ran into this spike, as it was digging the

hole, after progressing 6 inches or so downward. It did not seem able

to go around the spike, although it had enlarged the cavity an inch on

each side of the heavy nail and had cut away the wood for 2 inches or

more below the spike. But the job proved to be too much for the bird,

and it eventually gave up and disappeared. I told my friend to pull

the spike and maybe next year the bird would be back. He followed my

suggestion, and, the following spring, a Harris’s woodpecker showed up,

dug a hole in the stub, and raised its young. This was repeated the

next spring after, and then no return of the bird. So quite likely it

may have been the same woodpecker.”

_Eggs._--Harris’s woodpecker apparently lays four or five eggs,

probably seldom fewer or more. Bendire (1895) was evidently unable to

locate any properly identified eggs of this race, and I have not fared

much better. The eggs are probably indistinguishable from the eggs of

other hairy woodpeckers of similar size. The measurements of 34 eggs

average 25.29 by 18.91 millimeters; the eggs showing the four extremes

measure =27.9= by 19.6, 25.46 by =20.32=, =22.86= by 18.29, and 23.5 by

=17.5= millimeters.

_Food._--J. A. Neff (1928) says:

A total of 57 stomachs of hairy woodpeckers were taken for

the present study, over three fourths of them of the Harris

type. The months were represented by fairly even numbers of

specimens. Analysis of these stomachs shows a considerable

variation from the results of Professor Beal’s California

studies. The total animal food averaged 82.00 percent,

while vegetable matter made up the other 18.00 percent.

The larvae of wood-boring beetles, Cerambycids and

Buprestids, composed 49.00 percent of the total. This total

is unexceeded in Federal studies of birds. Since these two

groups of borers include species doing enormous damage to

both forest and ornamental trees, as well as to orchards,

this item of food alone almost settles the question of the

utility of having woodpeckers. * * *

The vegetable food was of little value economically.

Fruits, of small wild varieties, totaled 6.00 percent, and

seeds, mostly of coniferous trees, averaged 12.00 percent.

_Behavior._--Taylor and Shaw (1927) made the following interesting

observation:

As is well known, western Washington is a region of copious

rainfall. During the frequent downpours one can not help

speculating on the manner in which the different birds and

mammals avoid injury from the damp and chill of the storm.

The thick foliage of firs and hemlocks is well suited, in

many instances, to serve as a thatched roof: and in the

deep woods there are many big branches and large logs under

which birds--and mammals too--find dry retreats. During

the heaviest rain of the summer a Harris woodpecker was

frightened from its shelter beneath a huge log in the heavy

forest of Tahoma Creek Canyon. Here the bird was keeping

perfectly dry. One can imagine its displeasure at being

driven out from its comfortable refuge into the drenching

rain.

_Winter._--D. E. Brown says in his notes: “Early in fall Harris’s

woodpecker very often excavates a cavity that is its winter home. It

can be found there every night and quite often in the daytime on stormy

days. It is not always secure in this retreat. Such a cavity was made

in a partly dead stub, about 20 feet high, just back of my house.

Frequent visits were made to find out how much the nest was used. At

first the bird, a male, would fly to a nearby tree when I rapped on the

stub, but soon it contented itself with just coming to the opening.

One time, while I was in plain sight of the stub, a western pileated

woodpecker alighted at the cavity and proceeded to open it up, which it

did clear to the bottom in less than three minutes. It had taken the

Harris about a week to dig it out.”

Dawson and Bowles (1909) say: “The Harris woodpecker visits the

winter troupes only in a patronizing way. He is far too restless and

independent to be counted a constant member of any little gossip club,

and, except briefly during the mating season and in the family circle,

he is rarely to be seen in the company of his own kind.”

DRYOBATES VILLOSUS HYLOSCOPUS Cabanis and Heine

CABANIS’S WOODPECKER

HABITS

The hairy woodpeckers of the coast district of California from

Mendocino County southward, the mountains of southern California,

and the southern Sierra Nevada, as far east as White Mountains of

California, are now known by the above name. This race is somewhat

smaller than _harrisi_ and decidedly smaller than _orius_, its

neighboring races to the northward; its under parts are much lighter

colored than in _harrisi_; these parts are described by Ridgway

(1914) as “dull grayish or brownish white or pale drab-grayish or

buffy grayish.” This and the Sierra woodpecker (_orius_) seem to

form connecting links between the dark-breasted _harrisi_ and the

white-breasted Rocky Mountain forms, _monticola_ and _leucothorectis_,

both of which are decidedly larger also.

Dr. Joseph Grinnell (1908) says of the distribution of this woodpecker

in the San Bernardino Mountains in southern California: “This was the

most widely distributed species of woodpecker in the region, occurring

throughout the timbered portions, irrespective of zones. It was common

from Santa Ana Cañon to the summit of Sugarloaf, 9,800 feet, and nearly

to timber line on San Gorgonio peak. On the desert side the species was

noted as low as Cactus Flat, 6,000 feet, where one was seen in some

golden oaks in a ravine, August 16, 1905.”

_Courtship._--The drumming of woodpeckers in the spring on some

resonant limb or tree trunk is an important part of the courtship

urge, as a warning to any rival, or as a call to a possible mate. Dr.

Grinnell (1908) shows how the manner of drumming may also serve as

a recognition call; he says: “The resonant rattling drum identified

this species from any other of this region. Near Dry lake, 9,000

feet altitude, dead tamarack pines were selected for this purpose,

and on June 23, 1905, I listened for many minutes to a remarkable

demonstration of this kind. Different branches were tattoed in rapid

succession, so that a xylophone-like variety of tones was produced,

very impressive and far-carrying through the otherwise quiet forest.”

_Nesting._--Major Bendire (1895) writes:

In California Cabanis’ Woodpecker is common in the

mountains, but in the lowlands in the southern part of this

State Mr. F. Stephens considers it a rather rare summer

resident. He found it breeding in a cottonwood tree, near

San Bernardino, on March 21, 1885. Mr. Lyman Belding took

several nests of this subspecies in Calaveras County, in

the Sierra Nevadas; in one, found on June 6, 1879, which

had been excavated in a dead pine stump, 12 feet from the

ground, the eggs, three in number, were on the point of

hatching. In his notes he says: “I scared the female from

it and prevented her return by inserting a stick, the end

of which protruded for several feet. When she found she

could not enter she gave several cries, which brought

the male, who hopped up and down the stick a few times,

striking it with his bill and screaming angrily, pausing

occasionally, and apparently deliberating on the best

method of extracting it.” Another nest, found by him on

July 10, 1880, was located only 3 feet from the ground, and

contained young which were still in the nest on the 20th.

Mr. Charles A. Allen informs me that along the Sacramento

River, in California, it breeds in sycamores and willows,

but that it is not common there.

_Eggs._--Major Bendire (1895) says:

The number of eggs laid to a set varies from three to six;

those of four are by far the most common; sets of five are

only occasionally met with, while sets of six are very

unusual. * * * The eggs lie on the fine chips left in the

bottom of the cavity, and are occasionally well packed into

these, so that only about one-half of the egg is visible.

They resemble the eggs of _Dryobates villosus_ in color,

but those of an elliptical ovate shape are more common than

the oval and elliptical ovals, averaging, therefore, more

in length, while there is proportionally less difference in

their short diameter.

The measurements of 23 eggs average 24.49 by 18.38 millimeters; the

eggs showing the four extremes measure =25.7= by 18.2, 24.2 by =19.7=,

=22.8= by 18.1, and 24.9 by =16.5= millimeters.

_Young._--Mrs. Irene G. Wheelock (1904) says: “Both sexes share the

labors of excavating, brooding the eggs, and feeding the young.

Incubation lasts about fifteen days, and the young remain nearly four

weeks in the nest, being fed most of that time by regurgitation. After

leaving they are fed by the parents for at least two weeks, and usually

return to the nest at night to sleep.”

_Food._--W. Leon Dawson (1923) writes: “Nearly half of the Cabanis

Woodpecker’s food consists of the larvae of wood-boring beetles

(the _Cerambycidae_ and _Buprestidae_); and of the remainder the

caterpillars of various injurious moths form a large per cent. Wild

raspberries and blackberries are eaten in summer, and certain hardy

fruits, such as cornel berries, acorns, and the pits of the islay, or

evergreen cherry (_Prunus ilicifolia_), eke out the winter sustenance.”

Referring to its manner of feeding, Milton P. Skinner says in his

notes: “On August 10, 1933, I saw a Cabanis working on both the trunk

and the limbs of a small Douglas fir. It worked all around a horizontal

limb and really seemed to be under the limb more than above. It also

worked on upright branches as well. I have also seen a Cabanis feeding

on the bark of a lodgepole pine. One day I found one on a dead black

oak, scaling off dead bark to get at the insects beneath. So far as I

can tell, these birds, in the Sequoia National Park, seem to prefer to

pick food from the surface and furrows in the bark, and do not bore

into the bark and wood as much as other woodpeckers. During my work

among the Big Trees, I noticed that these birds seem to avoid the

sequoia’s bark; but at one place I found a living tree with many holes

bored in the old wood of its charred base, where it was unprotected by

bark.”

_Behavior._--Mr. Skinner’s notes say that “this woodpecker has quite a

few mannerisms of its own. One, seen flying across a meadow, went first

to the limbs of Douglas firs, then to a small dead limb of a sequoia,

then to the limb of a fir, and then to the trunk of the same fir. It

perched lengthwise of limb and trunk each time. And this procedure was

followed again and again on different days. Usually the Cabanis perches

crosswise on a horizontal limb, especially when resting or preening,

but lengthwise on erect, or nearly erect, trunks and limbs when feeding.

“Although this woodpecker almost always flies to the exact spot it

selects, its flight through the forest is undulatory. The undulations

are due to the fact that it progresses by a series of wing beats. At

the end of each series, it seems to actually close its wings and shoot

forward with the impetus gained.”

DRYOBATES VILLOSUS MONTICOLA Anthony

ROCKY MOUNTAIN HAIRY WOODPECKER

HABITS

This large, white-breasted hairy woodpecker inhabits the Rocky Mountain

region, in the Canadian and Transition Zones, from central British

Columbia and Montana southward to eastern Utah and northern New Mexico,

and eastward to western South Dakota and western Nebraska. Ridgway

(1914) characterizes it as “similar, in large size and whiteness of

under parts, to _D. v. septentrionalis_, but with white spots on

wing-coverts much reduced in size or number, or altogether wanting.” It

evidently intergrades with _septentrionalis_ in Montana and Wyoming

and probably with the more western races west of the Rocky Mountains.

Milton P. Skinner tells me that in the Yellowstone National Park it

“occurs at all elevations from the lowest at 5,500 feet to timberline

at 9,500 feet above sea level, but never far from a tree of some kind.

It is a resident bird here but moves down from the mountain heights at

the approach of winter.”

Aretas A. Saunders (1921) says of its status in Montana: “A common

permanent resident throughout the western half of the state in the

mountains. Winters mainly in the valleys in cottonwood groves, but does

not breed there. * * * The eastern limits of its range are evidently

in the eastern foothills of the mountains. Just what form breeds in

the more eastern mountain ranges is not definitely determined. In the

mountains this bird has been recorded by all observers. It is common

everywhere, and usually the commonest of the mountain woodpeckers.”

_Nesting._--The following remarks by Major Bendire (1895), under

_hyloscopus_, evidently refer to this subspecies: “Mr. Denis Gale found

it breeding in Boulder County, Colorado, on May 28, 1886, in a live

aspen tree, at an altitude of about 8,500 feet. The nest contained five

eggs, in which incubation was somewhat advanced. Mr. William G. Smith

also reports it as common in Colorado, coming down into the valleys in

winter. He says it is the earliest of the Woodpeckers to breed, that it

commences nesting in the latter part of April, and usually excavates

its holes in old dead pines, frequently at a considerable distance from

the ground, and that he has seen full-grown young by June 1.”

J. K. Jensen (1923) says of this woodpecker, in northern Santa Fe

County, N. Mex.:

Quite common in the Sangre de Cristo Mountains, from 8,000

to 11,000 feet. June 21, 1920, I found a nest thirty feet

up in a large quaking aspen. This tree stands on the edge

of a place where an avalanche has plowed its way down

through the timber on the mountain side, depositing trees

and rocks in a great heap for hundreds of feet around the

tree. The nest contained young, and judging from the noise

they made, were quite well developed. The parent birds were

very noisy.

May 22, 1921, I made my way through four feet of snow to

the same tree. A new nest had been made, and the female

flew off when I was about 150 feet away. I cut into the

nest and found a set of four eggs on which incubation had

just commenced. The altitude at this point is 11,000 feet.

May 26, 1922, I found a nest with young about seventy-five

feet up in an aspen. This was in Santa Fe Canyon at an

altitude of 8,000 feet.

_Eggs._--The eggs of the Rocky Mountain hairy woodpecker are similar to

the eggs of other hairy woodpeckers of similar size. The measurements

of 33 eggs average 24.89 by 18.49 millimeters; the eggs showing the

four extremes measure =28.08= by 18.03, 27.0 by =20.1=, =23.37= by

17.78, and 24.38 by =17.27= millimeters.

_Food._--Mr. Skinner says, in his notes, that this woodpecker “seeks

its food on the trunks of lodgepole and _flexilis_ pines, cedars,

firs, aspens, willows, and even electric-light and telephone poles; it

prefers dead and diseased trees and stubs to work on, probably because

of more borers and grubs. At Basin, and over 7,000 feet elevation,

I found a female where I could watch her, only 5 feet away from the

lodgepole trunk on which she was working. She worked down, tapping

here and there as she went. Whenever a tap revealed a borer, she

scaled off the bark with quick right and left strokes, having a slight

lever motion at the end, and always secured from one to six bark-borer

grubs. Evidently the tap told her whether it was worth while to search

further, for she never made a mistake and performed no useless labor.”

J. A. Munro (1930) writes: “During the winter of 1928-29 a male hairy

woodpecker frequently was seen feeding on Virginia creeper berries in

competition with several red-shafted flickers. On one occasion the same

bird visited an apple tree, attracted by a few apples that still clung

to the bare branches. Standing crossways on a branch, in the ordinary

position of a perching bird, he rapidly stabbed his bill downward into

the top of an apple. After doing this several times he flew to another

portion of the tree and repeated the performance.”

DRYOBATES VILLOSUS PICOIDEUS Osgood

QUEEN CHARLOTTE WOODPECKER

HABITS

Life histories of North American woodpeckers · 第 4 / 43 段