The second section of the order GLIRES, containing the following
families--those that are not Indian being in italics:--
_Myoxidiae_, _Lophiomyidae_, Muridae, Spalacidae, _Geomyidae_,
_Theridomyidae_ (fossil), Dipodidae.
The molar dentition is from 3--3/3--3 to 6--6/6--6, the former being
the usual number; the tibia and fibula are united for at least a third
of their length; the zygomatic arch is slender, and the malar process
rarely extends so far forwards as in the preceding section, and is
generally supported below by a continuation of the maxillary
zygomatic process; the collar-bones are perfect (except in
_Lophiomyidae_). Upper lip cleft; the muffle small and naked; tail
cylindrical, sometimes hairy, but commonly covered with scales
arranged in rings.
In all the Indian mammalogy this section is probably the most
difficult to write about. Our knowledge of the smaller rodents is
extremely imperfect, and is just engaging increased attention. In
the meanwhile I feel that, while I make use of such material as is
now available, before long much will have to be revised and corrected
after the exhaustive inquiries now being made by Dr. Anderson are
published.
The Indian families with which we have to deal are but three--the
_Muridae_, _Spalacidae_, and the _Dipodidae_. The _Arvicolidae_ of
Jerdon's work is merely a sub-family of _Muridae_. Of these the
_Muridae_ take the first place, as containing the greater number of
genera. It is estimated that the total number of species known of
this family throughout the world exceed 330, of which probably not
more than one-fourth or fifth are to be found in India and adjacent
countries.
FAMILY MURIDAE.
CHARACTER.--"Lower incisors compressed; no premolars; molars rooted
or rootless, tuberculate or with angular enamel folds; frontals
contracted; infra-orbital opening in typical forms high,
perpendicular, wide above and narrowed below, with the lower root
of the maxillary zygomatic process more or less flattened into a
perpendicular plate; very rarely the opening is either large and oval,
or small and sub-triangular. Malar short and slender, generally
reduced to a splint between the maxillary and squamosal processes;
external characters very variable; pollex rudimentary, but often
with a small nail; tail generally sub-naked and scaly, rarely densely
haired."--_Alston_, 'P. Z. S.' 1876.
This family is divided into about ten sub-families, of which the
Indian ones are as follows: _Platacanthyominae_; _Gerbillinae_;
_Phlaeomyinae_; _Murinae_; _Arvicolinae_; _Cricetinae_.
The other four are _Sminthinae_, _Hydromyinae_, _Dendromyinae_, and
_Siphneinae_, none of which are found within our limits.
_GENUS PLATACANTHOMYS_.
CHARACTER.--Molars 3/3, divided into transverse laminae;
infra-orbital opening as in typical _Muridae_; incisive foramina and
auditory bullae small; form _myoxine_ (or dormouse-like); fur mixed
with flat spines; tail densely hairy. The general resemblance of this
animal to the dormouse (_Myoxus_) is striking, to which its hairy
tail and its habits conduce, but on closer examination its small eyes,
thin ears, short thumb of the fore-foot bring it into the murine
family. The genus was first noted and named by Blyth, who seemed
inclined to class it as a dormouse, but this has not been upheld for
the reasons given above, and also that _Platacanthomys_ has the
normal _murine_ number of molars, viz.: 3--3/3--3, whereas _Myoxus_
has an additional premolar above and below. These points were first
brought to notice by Prof. Peters of Berlin (_see_ 'P. Z. S.' 1865,
p. 397). There is a coloured plate of the animal in the same volume,
but it is not so well executed as most of the illustrations in the
Society's works.
NO. 316. PLATACANTHOMYS LASIURUS.
_The Long-tailed Spiny Mouse_ (_Jerdon's No. 198_).
HABITAT.--Southern India.
DESCRIPTION.--Light rufescent brown; the under fur paler, more
rufous on the forehead and crown; whiskers black; under parts dull
white; the hairs on the tail, which are arranged distichously, are
darker than those of the body, infuscated except at the tip of the
tail, where they are whitish; the muzzle is acute; ears moderate and
naked; the fur above is mixed densely with sharp flat spines; the
under coat is delicate and fine; the few spines on the lower parts
are smaller and finer; the thumb is without a nail.
SIZE.--Head and body, 6 inches; tail, 3-1/2, or five inches including
the hair; planta, 1 inch.
This species was discovered by the Rev. Mr. Baker in the Western Ghats
of Malabar, and in Cochin and Travancore, at an elevation of about
3000 feet. He writes of it: "It lives in clefts in the rocks and hollow
trees, and is said to hoard ears of grain and roots, seldom comes
into the native huts, and in that particular neighbourhood the
hillmen told me they are very numerous. I know they are to be found
in the rocky mountains of Travancore, but I have never met with them
on the plains." In another place he adds: "I have been spending the
last three weeks in the Ghats, and, amongst other things, had a great
hunt for the new spiny dormice. They are most abundant, I find, in
the elevated vales and ravines, living only in the magnificent old
trees there, in which they hollow out little cavities, filling them
with leaves and moss. The hill people call them the 'pepper-rat,'
from their destroying large quantities of ripe pepper (_Piper
nigrum_). Angely and jackfruit (_Artocarpus ovalifolia_ and
_integrifolia_) are much subject to their ravages. Large numbers of
the _shunda_ palm (_Caryota_) are found in these hills, and toddy
is collected from them. These dormice eat through the covering of
the pot as suspended, and enjoy themselves. Two were brought to me
in the pots half drowned. I procured in one morning sixteen specimens.
The method employed in obtaining them was to tie long bamboos (with
thin little branches left on them to climb by) to the trees; and,
when the hole was reached, the man cut the entrance large enough to
admit his hand, and took out the nest with the animals rolled up in
it, put the whole into a bag made of bark, and brought it down. They
actually reached the bottom sometimes without being disturbed. It
was very wet, cold weather, and they may have been somewhat torpid;
but I started a large brown rat at the foot of one of these trees,
which ran up the stem into a hole, and four dormice were out in a
minute from it, apparently in terror of their large friend. There
were no traces of hoarding in any of the holes, but the soft bark
of the trees was a good deal gnawed in places. I had two of these
dormice alive for some time, but, as they bit and gnawed at everything
intended to keep them in durance, I was obliged to kill both. I
noticed that when their tails were elevated, the hairs were perfectly
erect like a bottle-brush" ('Proc. As. Soc. Beng.' 1859, p. 290).
SUB-FAMILY GERBILLINAE.
Incisors narrow; molars divided into transverse laminae; pterygoid
fossae short; auditory bullae usually large; hind limbs very long;
tail long and hairy.
_GENUS GERBILLUS_.
Form murine, with the exception of the elongated hind-limbs; muzzle
pointed; ears moderate and oval; eyes very large and bright;
occipital region broad; auditory bullae large; upper incisors
grooved; first molar with three laminae, the second with two, and
third with one only; hinder tarsus and toes much elongated; the
fore-limbs small; tail long and hairy, with a tuft at the end.
[Illustration: Dentition of _Gerbillus_ (magnified).]
NO. 317. GERBILLUS INDICUS.
_The Indian Jerboa-Rat, or Kangaroo-Rat_ (_Jerdon's No. 170_).
NATIVE NAMES.--_Hirna-mus_, Hindi; _Jhenku-indur_, Sanscrit and
Bengali; _Yeri-yelka_ of the Waddurs; _Tel-yelka_ of the Yanadees;
_Billa-ilei_, Canarese.
HABITAT.--All over India and in Ceylon, but apparently not in Burmah.
DESCRIPTION.--Light fulvous brown above or fawn colour, paling on
the sides; under-parts white; the hairs of the back are ashy at the
base, with fulvous tips, a few thin black hairs intermixed chiefly
on the side and cheeks.
The eyebrow is whitish; whiskers long and black and a few grey; the
nose is elongated; the upper jaw projecting nearly half an inch
beyond the lower; tail, which is longer than the body, is blackish
above and below, pale laterally, and terminates with a black tufted
tip; the ears are large and nearly naked; the eye is particularly
large and lustrous, which, with its graceful bounds, have given it
its Indian name of "antelope-rat" (_Hirna-mus_).
SIZE.--Head and body, about 7 inches; tail, 8-1/2 inches; fore-foot,
5/10 inch; hind-foot, 2 inches. Weight, 6-3/4 ounces.
This graceful little creature frequents bare plains and sandy
country in general, where it forms extensive burrows. Hardwicke
writes of it: "These animals are very numerous about cultivated lands,
and particularly destructive to wheat and barley crops, of which they
lay up considerable hoards in spacious burrows. A tribe of low-caste
Hindus, called Kunjers, go in quest of them at proper seasons to
plunder their hoards, and often within the space of twenty yards
square find as much corn in the ear as could be crammed in a bushel."
Sir Walter Elliot's account of their burrows is most interesting.
He says: "The entrances, which are numerous, are small, from which
the passage descends with a rapid slope for two or three feet, then
runs along horizontally, and sends off branches in different
directions. These galleries generally terminate in chambers from
half a foot to a foot in width, containing a bed of dried grass.
Sometimes one chamber communicates with another furnished in like
manner, whilst others appear to be deserted, and the entrances closed
with clay. The centre chamber in one burrow was very large, which
the Wuddurs attributed to its being the common apartment, and said
that the females occupied the smaller ones with their young. They
do not hoard their food, but issue from their burrows every evening,
and run and hop about, sitting on their hind legs to look round,
making astonishing leaps, and on the slightest alarm flying into
their holes." This account differs from that of Hardwicke as regards
the hoarding of food, and from what I can learn is the more correct.
The food of this animal is grain, grass, and roots, but Kellaart
mentions certain carnivorous propensities, for one night several of
them nearly devoured an albino rat which had been put into the same
cage with them. McMaster says of its agility: "I have seen them when
released from a trap baffle and elude dogs in the most extraordinary
manner by wonderful jumps made over the backs, and apparently into
the very teeth of their pursuers."
Buchanan-Hamilton's assertion that "these animals live in holes
which they dig in the abrupt banks of rivers and ponds" is misleading.
They may do so occasionally, but in general they choose sandy plains.
The female is prolific, bringing forth from eight to twelve young
ones, and Dr. Jerdon states that it is said to have occasionally as
many as sixteen to twenty. With regard to Kellaart's accusation of
its being carnivorous at times, I may say I have noticed such
tendencies amongst several other rodents which are supposed to be
purely vegetarians. I have also known ruminants take to flesh-eating
when opportunity offered.
NO. 318. GERBILLUS HURRIANAE.
_The Desert Jerboa-Rat_ (_Jerdon's No. 171_).
HABITAT.--The sandy deserts west of the Jumna and Hurriana; also in
Afghanistan according to Horsfield's Catalogue, and probably in
Rajpootana, Sindh, and the Punjab.
DESCRIPTION.--Pale rufous or sandy above, with fine dusky lines, the
hairs being blackish at the base, the rest fawn coloured, with a
blackish tip very minute; sides paler, with fewer dusky lines;
under-parts white, tinged more or less with fulvous or fawn on the
belly; limbs pale fawn; orbits pale; whiskers whitish, a few of the
upper ones dark; tail yellowish-rufous or fawn colour throughout,
with a line of dusky brown hairs on the upper surface of the terminal
half, gradually increasing in length to the tips.
SIZE.--Smaller than the last species. Head and body, 5 inches; tail,
4-1/2.
Jerdon says of this rat that it is "exceedingly numerous in the sandy
downs and sand-hills of Hurriana, both in jungles and in bare plains,
especially in the former, and a colony may be seen at the foot of
every large shrub almost. I found that it had been feeding on the
kernel of the nut of the common _Salvadora oleifolia_, gnawing
through the hard nut and extracting the whole of the kernel. Unlike
the last species, this rat, during the cold weather at all events,
is very generally seen outside its holes at all hours, scuttling in
on the near approach of any one, but soon cautiously popping its head
out of its hole and again issuing forth. In the localities it
frequents it is far more abundant than I have ever seen _G. Indicus_
in the most favourable spots" ('Mammals of India,' p. 186).
NO. 319. GERBILLUS CRYPTORHINUS.
_The Lobe-nosed Jerboa-Rat_.
HABITAT.--Yarkand.
DESCRIPTION.--after Mr. Blanford, who first described and named the
species: "Colour above sandy rufescent, some specimens rather more
rufous than others; below white, the two colours sharply divided on
the sides; cheeks pale; supercilia whitish; feet white; tail above
rather more rufous than the back, paler and occasionally whitish
below, becoming dark brown or blackish above near the end, and with
the slight tuft of longer hairs at the end of the same dark colour;
fur soft and glossy, about half an inch long in the middle of the
back, all the basal portion being at least three-quarters of the
length, dark ashy; the terminal portion pale yellow brown to pale
rufous, with numerous longer hairs with black tips mixed; on the
under surface the hairs are white throughout; on the tail the hair
is rather short, coarse, and close together; there are a very few
longer black tips mixed, but scarcely enough to produce an effect
in the general colour.
"The ears are oval and of moderate length; densely clad with brown
hairs on the anterior portion of the outer surface, and with a fringe
of longer hairs on the anterior margin; the posterior portion of the
external surface is nearly naked, except near the margin, and the
anterior portion of the inner surface is completely destitute of hair,
but the inner surface is more hairy near the hinder margin. The
whiskers are very numerous, the longest slightly exceeding the head;
the uppermost behind being black, all the rest white; all are mixed
at the base with long hairs, which cover the side of the nose; soles
of the fore-feet with scattered white hairs, but nearly naked; those
of the hind-feet densely covered with hair everywhere except at the
extreme tips of the toes and at the heel.
"Mammae, eight--four pectoral and four inguinal, as usual in the
genus.
"The most remarkable character of these species is the presence at
the end of the snout of a semi-circular lobe, which forms a flap
completely covering the openings of the nostrils. This lobe can, of
course, only be well seen in the specimens preserved in spirit. In
the dried skin its presence can sometimes be detected, but not always.
In the only spirit specimen, an adult female, the flap measures about
0.3 inch in breadth, and is barely an eighth of an inch long.
"It is hairy both outside and inside, the hairs being very short and
rather scattered inside; the surface below the nostrils covered by
the flap is also hairy. The use of this lobe is evidently to keep
out sand and dust from the air passages" (W. T. Blanford's 'Mammalia
of the Second Yarkand Mission,' p. 56).
SIZE.--Head and body, about 5-1/2 inches; tail, 5 inches; length of
fore-foot, 0.5 inch; hind-foot, 1.4 inch.
The peculiarity of the lobe, which was first detected by Mr. Oscar
Fraser in removing a skull from a spirit specimen, distinguishes this
species from the other Asiatic forms. There is also a peculiarity
in the skull noticed by Mr. Blanford, which is that the lachrymal
process, instead of being anchylosed to the adjoining bones, as in
others of the genus, is free, and this species is therefore
distinguished from the one most resembling it, _G. unguiculatus_
from Chinese Mongolia, in which the lachrymal process is united to
the frontal.
NO. 320. GERBILLUS ERYTHRURUS.
_The Red-tailed Jerboa-Rat_.
HABITAT.--Afghanistan and Persia.
DESCRIPTION.--Rufous brown above, with a few long black hairs, more
numerous on the rump and thighs; under fur slaty; under-parts white,
gradually blending with the colour of the sides; ears much larger
than in the last species, hairy outside and near the margin inside;
soles of hind feet and toes thickly covered with hair, except on the
hinder half of the tarsus; tail very rufous--brown with a black tip,
black hairs are scattered along the upper surface, and form a black
band towards the end above, finally covering the whole tip.
SIZE.--Head and body, about 6 inches; tail, equal.
Mr. Blanford, to whose 'Eastern Persia' I am chiefly indebted for
the above description, writes: "From _G. Hurrianae_, which Jerdon
thought might probably be the same, the present form is distinguished
by its much larger ears and by the hind feet, and especially the toes,
being more thickly covered with hair beneath; the fur too is longer
and the colour browner on the back; the tail is more rufous, and the
tip blacker; the skull is larger and broader; the nasal portion more
elongate and less concave above, and the hind upper molar has a
distinct talon, or rudimentary second transverse ridge, in young
specimens, traces of which may be detected in the form of the worn
tooth."
Its habits are similar to those of the last species.
NO. 321. GERBILLUS NANUS.
_The Dwarf Jerboa-Rat_.
HABITAT.--Baluchistan.
DESCRIPTION.--The fur is soft and long, rufous brown or fawn colour
above, white below, the colours being less sharply distinguished
than in _G. Indicus_; the hairs of the upper parts have no black tips,
and the basal two-thirds are slaty grey. There is a broad white
supercilium in front, joining the white area of the sides of the face,
so that the brown of the nose is reduced to a rather narrow band;
ears almost naked, a few short whitish hairs near the edge only;
whiskers nearly all white; a few of the upper hairs brown near the
base; feet white above, naked beneath, tail light brown above,
whitish beneath; towards the end a band of darker brown hairs runs
along the upper portion, those at the end lengthened; but there is
a less marked tuft than usual, and there are no black hairs at the
end (Blanford's 'Eastern Persia,' vol. ii. p. 72, _with plate_).
SIZE.--Head and body, 2.6 inches; tail, exclusive of hair, 4.5
inches; hair, 0.55 inches.
This curious little animal was first found and named by Mr. W. T.
Blanford, who obtained two specimens, with others of _G. Hurrianae_,
in a large area of ground that was flooded. He at first supposed them
to be the young of _G. Indicus_, but found on subsequent examination
that they were full grown.
SUB-FAMILY PHLOEMYINAE.
Incisors broad; molars divided into transverse laminae;
infra-orbital opening typical; claws large.
_GENUS NESOKIA_.
Muzzle blunt; ears moderate; claws long; fur rather harsh; tail short,
scaly, sparsely haired; palate narrow; incisive foramina short;
auditory bullae rather small; incisors broad; first molars with
three laminae, the rest with only two.--_Alston_.
There has been some confusion regarding the species of this genus.
Jerdon, in his 'Mammals of India,' gives only two, including
_Arvicola Indica_ and _Mus kok_ of Gray, _Mus providens_ of Elliot,
and _Mus pyctoris_ of Hodgson, under _Nesokia Indica_, and
classifying _Nesokia Huttoni_ with _N. Hardwickii_; but Dr. Anderson,
after a most careful examination of specimens from all parts of India,
has proved the distinctness of _Mus providens vel kok_ from the
species called by Jerdon _Nesokia Indica_, which, being a synonym
of _N. Hardwickii_, he has now renamed _Mus (Nesokia) Blythianus_
(_see_ 'Jour. As. Soc. Beng.' 1878, vol. xlvii. pt. ii.), and Mr.
Blanford had clearly demonstrated that _N. Huttoni_ is a distinct
species from _N. Hardwickii_ ('Zool. of Persia,' vol. ii. p. 59).
NO. 322. NESOKIA HARDWICKII.
_Hardwick's Field-Rat_ (_Jerdon's No. 173_).
HABITAT.--North-western India.
DESCRIPTION.--General colour sandy brown on the upper parts, paler
on the sides, dusky grey, with a tinge of yellowish-rufous on the
under-parts; muzzle, feet, and tail flesh-coloured; ears of the same,
but rather darker; head short and bluff; muzzle broad and deep; eye
moderately large; ears moderate, rounded, clad with minute hairs;
fur soft and moderately long, of three kinds, viz. short under-fur,
ordinary hairs, and mixed with them, especially on the back and rump,
numerous long black hairs which project a good way beyond the fur.
SIZE.--Head and body, nearly 8 inches; tail, about 4-1/2 inches.
It is probable that this species is identical with _Mus Griffithi_,
though the dimensions given by Horsfield ('Cat. Mam. Mus. E. I.
Comp.') and the description do not quite agree. He gives the size
of head and body at 6-1/2 inches; tail, 3 inches, and says that the
teeth are nearly white.
NO. 323. NESOKIA HUTTONI.
_Hutton's Field-Rat_.
HABITAT.--Northern India, Afghanistan and Persia.
DESCRIPTION.--Colour above from ferruginous brown to sandy brown,
lower parts isabelline, but frequently appear dark in consequence
of the fur being thin or worn; the basal portion dark slaty grey both
above and below the animal; hairs on the back soft and of moderate
length, a very few black hairs being scattered amongst the brown
ones; tail naked, and ears almost naked, the latter having only a
few extremely short hairs, thinly scattered, and the feet are covered
above very sparsely with short whitish hairs (_see_ Blanford's
'Persia,' vol. ii., for description and plate). Nose and feet
flesh-coloured; ears and tail darker and brownish; mammae eight, as
usual in the genus.
According to Dr. A. Barclay (quoted by Dr. Anderson) the holes of
this rat do not run deep, but ramify horizontally just below the
surface of the ground. It throws out a mound of earth at the exit
of the hole.
NO. 324. NESOKIA SCULLYI.
_Scully's Field-Rat_.
NATIVE NAME.--_Mughi_, Turki.
HABITAT.--Kashgaria at Sanju, south-east of Yarkand.
DESCRIPTION.--Light rufescent brown above, dirty white beneath;
fur fine and silky, blackish-grey at the base, and for two-thirds,
the last third of the longer hairs being fawn colour; face earthy
brown; whiskers black, tipped with white; ears very short,
semi-nude; feet and claws flesh-coloured; tail naked, with a few
scattered fine short hairs.
SIZE.--Head and body, 6.6 inches; tail, 5.2 inches.
NO. 325. NESOKIA PROVIDENS.
_The Southern India Field-Rat_ (_Jerdon's No. 172_).
NATIVE NAMES.--_Kok_, Canarese; _Golatta-koku_, Telegu of the
Yanadees; _Yea-kwet_ (?) Burmese.
HABITAT.--Southern India and Ceylon, probably Burmah, as one species
is mentioned there by Blyth.
DESCRIPTION.--Head short and truncated, with a deep muzzle; ears
nearly round, semi-nude, sparsely covered with minute hairs; eyes
moderately large, half-way between snout and ear; feet largish;
claws short and stout; tail nearly equalling length of head and body,
semi-nude, ringed, and with short brown bristly hairs round the
margin of the annuli; whiskers full and long; colour of the
fur--which is harsh and long, as in the rest of the genus, and of
the usual three kinds--is a brown, mixed with a tinge of fawn; the
under-parts are whitish, with a yellowish tinge; the nose, ears, and
feet are dark flesh-coloured or brownish, and the feet are covered
with short brown hair. The incisors are orange yellow; the claws
yellowish.
Sir Walter Elliot states that a variety found in red soil is much
redder in colour than that inhabiting the black land. The skull is
considerably smaller, according to Dr. Anderson, than that of the
Bengal _Nesokia_, _N. Blythiana_, of the same age, from which it is
also distinguished by its more outwardly arched malar process of the
maxillary, by its considerably smaller teeth and long but less open
anterior palatine foramina. The brain case is also relatively
shorter and more globular than that of _Nesokia Blythiana_.
SIZE.--Head and body, about 7 inches; tail, 6-1/2 inches.
The habits of this rat are similar to those of the Bengal species,
to which I will allude further on, and it has the same way of taking
to water when pursued.
Jerdon says that this rat is most destructive to tea-trees, biting
the roots just below the surface, more, he believes, because they
happen to come in the way of their burrows than to feed on them.
Sir Walter Elliot writes: "In its habits it is solitary, fierce,
living secluded in spacious burrows, in which it stores up large
quantities of grain during the harvest, and when that is consumed
lives upon the _huryale_ grass and other roots. The female produces
from eight to ten at a birth, which she sends out of her burrow as
soon as they are able to provide for themselves. When irritated it
utters a low grunting cry, like the bandicoot. The race of people
known by the name of Wuddurs, or tank-diggers, capture this animal
in great numbers as an article of food, and during the harvest they
plunder their earths of the grain stored up for their winter
consumption, which in favourable localities they find in such
quantities as to subsist almost entirely upon it during that season
of the year. A single burrow will sometimes yield as much as half
a seer (1 lb.) of grain, containing even whole ears of jowaree
(_Holchus sorghum_)." Sir Walter Elliot goes on to give a most
interesting account of the construction of the burrows of this
animal.
NO. 326. NESOKIA BLYTHIANA.
_The Bengal Field-Rat_.
NATIVE NAME.--_Yenkrai_, Bengalee.
HABITAT.--From Ghazipur in the North-west to Eastern Bengal and
Cachar. Very common about Calcutta.
DESCRIPTION.--Fur coarse as in the genus, profusely intermixed with
long piles, more numerous on the lumbar and sacral regions, which
project a long way beyond the ordinary pelage. The general colour
a dark brown with yellowish hairs intermingled, which give a somewhat
rufous tinge, paler beneath. Nose, ears, and feet flesh-coloured;
tail naked, ringed, and sparsely covered with short bristly hairs
at the margin of the rings; feet moderately large; claws short and
stout; eyes moderately large, placed a little nearer to the ear than
to the snout; ears rounded, semi-nude, covered with a fine down;
whiskers black; incisor teeth rich orange, but generally white
towards their tips.
The female has eight pairs of mammae.
SIZE.--Head and body, 8-1/4 inches; tail, 6-1/2 inches.
I have already alluded to the distinguishing features of the skull
of this species, as compared with _Nesokia providens_. From the skull
of _N. Hardwickii_ it differs in its considerably narrower incisors
and smaller and more irregularly laminated molars, and by its long
and open anterior palatine foramina. It has also a more arched skull
(_Anderson_).
This animal, which is included in Jerdon's _Nesokia Indica_, is very
generally distributed over Lower Bengal. In the neighbourhood of
Calcutta, Alipore for instance, it is abundant, and is a great
nuisance in gardens. It burrows in tortuous directions, only a few
inches below the ground, there being no definite plan, some being
more complicated than others--the principal passage leading to a
chamber containing a nest of leaves and grass. I have been told by
natives that large quantities of grain are stored by these rats. When
I first heard of its aquatic powers, I was led to believe that it
was a species of vole, and was particularly desirous to get one, not
being aware of any true water-rat in India. However, the reports of
the natives have been confirmed by what Sir Walter Elliot states
regarding the habits of _N. providens_, and by Dr. Anderson, who made
several experiments with these rats in captivity. He says: "To test
this aquatic power, I had two rats placed in a large wire birdcage,
and the cage partially submerged; if the rats, when in those
circumstances, were much annoyed, they immediately dived to the
bottom of the cage, where they could be observed running about under
water. I also had them removed from the cage, and let loose in the
large sheet of water in the Zoological Gardens, between the two iron
bridges. When let loose at the bank, and an attempt was made to catch
them, they immediately dived; and the stronger of the two did not
appear at the surface for some time, when it was observed at a
considerable distance from the bank making for the opposite side."
In confinement these rats are not engaging pets; they show a
considerable amount of surliness and ferocity. I have noticed that
on approaching the bars of the cage, one would grind its teeth, put
back its ears, and fly at you with a grunt.
NO. 327. NESOKIA BARCLAYIANA.
_Barclay's Field-Rat_.
HABITAT.--Northern India, the North-west and some parts of Bengal
(Purneah) and Assam.
DESCRIPTION.--General colour brownish; under surface silvery grey;
feet and muzzle flesh-colour; tail nearly black; claws horny white;
a white band from the nose through the eye; muzzle short and bluff;
forehead slightly arched; tail exceeding the length of the trunk,
but not equal to head and body, ringed, and sparsely clad; fur coarse;
piles moderately long.
SIZE.--Head and body, about 8-3/4 inches; tail, 7-1/4 inches.
This rat was first discovered by Dr. Arthur Barclay at Goona in
Central India, and apparently it appears to be identical with
specimens collected at Srinagar in Kashmir, in the Purneah district,
and in Cachar.
* * * * *
The next two have usually been classed as true _Mus_, and the latter
is to be found in Jerdon; but, from the breadth of the incisors and
the lamination of the molars, which are less sinuous and relatively
larger than in _Mus_, and from other characteristics of the skull,
they are nearer allied to _Nesokia_ than to the true rats.
NO. 328. MUS (NESOKIA) ELLIOTANUS.
_Elliot's Field-Rat_.
HABITAT.--Bengal, Assam, Khasia hills.
DESCRIPTION.--This rat is thus described by Dr. Anderson. It is the
nearest approach in size to the bandicoot: "Head short and deep;
muzzle deep and broad; eye half-way between ear and nose, moderately
large; ears not large, rounded, sparsely covered with short hairs;
feet large and well developed, with strong claws, and sparsely clad;
tail sparsely covered with short bristles on the margins of the
annuli, and nearly equalling the length of the body and head. Pelage
coarse, with moderately large piles, most numerous on the back;
vibrissae moderately long.
"General colour, above brown, with intermixed yellowish or pale
brown hairs producing much the same colour as in _M. (N.)
Blythianus_; paler on the sides, and passing into greyish on the
under-parts; nose and feet flesh-coloured; ears dark brown; tail
blackish" ('J. A. S. B.' 1878, vol. xlvii; pt. ii. p. 231).
NO. 329. MUS (NESOKIA) GIGANTEUS.
_The Bandicoot_ (_Jerdon's No. 174_).
NATIVE NAMES.--_Indur_, Sanscrit; _Ghunse_, Hindi; _Ikria_,
Bengali; _Heggin_, Canarese; _Pandi-koku_, i.e. pig-rat, Telegu;
_Oora-meyoo_, Singhalese.
HABITAT.--Throughout India; also in Ceylon.
DESCRIPTION.--Fur coarse, consisting of the three kinds, of which
the coarser piles are very long, and almost hide the general pelage
on the lumbar and dorsal regions. These piles are almost absent on
the head, neck, and sides; general colour earthy brown, with
yellowish hairs intermixed; the piles blackish-brown; under-parts
dusky brown, mixed with grey; limbs brownish; nose, inside of ear
and feet flesh-coloured; tail black, ringed, and sparsely haired.
The female has twelve mammae.
SIZE.--Head and body, from 12 to 15 inches; tail, from 11 to 13.
Weight, about 3 lbs.
This is a well known rat, but it is not common in Calcutta, although
supposed to be so. People frequently mistake very large specimens
of the common brown house-rat (_Mus decumanus_) for this animal,
which, Blyth remarks, is rare here. Jerdon states that it is common
in the fort of Madras, where he killed many, some of large size. When
assailed it grunts like a pig, hence its Telegu name _Pandi-koku_,
from which the word bandicoot is derived. McMaster states that the
bandicoot, though so formidable in appearance, does not show so good
a fight as an ordinary English rat, being a sluggish and cowardly
animal; and though, from its size and weight, it takes a good deal
of worrying, it seldom does much in self defence, and any moderately
good dog can kill it with ease. It is however a most destructive
animal, doing much damage to granaries, gardens, and even
poultry-yards. In some parts of the country, as for instance Fort
St. George in Madras, Government used to pay a reward of one anna
for every bandicoot killed within the walls.
SUB-FAMILY CRICETINAE.
CHARACTER.--Molars tuberculate; infra-orbital opening sub-typical,
not much narrowed below, and the perpendicular plate little
developed; large internal cheek pouches.--_Alston_.
_GENUS CRICETUS--THE HAMSTERS_.
Form thick-set, with short limbs and tail, the latter sparsely haired,
not scaly. "Skull with marked but rounded supra-orbital ridges
continued into temporal ridges; coronoid process high and falcate"
(_Alston_). The incisors are plain; the molars tuberculated when
young, but in the old animal the tubercles are worn down and exhibit
laminae. They are very nearly related to the true rats, but differ
conspicuously in the possession of large cheek pouches--like those
of the pouched monkeys, into which they stuff the grain they carry
to their burrows. The hind-limbs have five toes, the fore-feet four
only, the thumb being represented by a wart. The European hamster
is a very destructive little animal, from its numbers and the
quantity of grain it stores away in its burrows. They have two sets
of burrows for summer and winter, the latter being the deepest and
most complicated. They pass the winter in a torpid state, but make
up for it by their activity in the summer months. The young are
produced twice in the year and in number varying from six to eighteen,
and they develop very rapidly. Their eyes open in about a week, and
when a fortnight old the parents drive them off to shift for
themselves. The European hamster is a most savage little creature,
and has been known to attack even a red-hot bar, and hold on in spite
of the pain.
[Illustration: Dentition of _Cricetus_.]
[Illustration: _Cricetus_.]
* * * * *
The two following are dwarf species--_Cricetulus_ of some
authors:--[22]
[Footnote 22: Dallas mentions (Cassell's 'Nat. Hist.') a species
from Kumaon, _Cricetus songarus_.]
NO. 330. CRICETUS PHAEUS.
_The Persian Hamster_.
HABITAT.--Yarkand, Gilgit, Persia.
DESCRIPTION.--Cinereous above, white below; the colour varies from
pure ashy grey to grey with an isabelline tinge.--_Blanford_.
SIZE.--Head and body, about 4 inches; tail, 1-1/4 inches.
NO. 331. CRICETUS FULVUS.
_The Sandy Hamster_.
HABITAT.--Yarkand, Gilgit.
DESCRIPTION.--Colour above light sandy brown to sandy grey; no band
down the back; lower parts, feet, and tail white; fur very soft, fully
half an inch long in the middle of the back in some specimens. Rather
larger than the last species. (_See_ Blanford's 'Second Yarkand
Mission,' p. 45.)
SIZE.--Head and body about 4-1/2 inches; tail about 1-1/2 inches.
SUB-FAMILY MURINAE.
CHARACTER.--Molars tuberculate, at least in youth; infra-orbital
opening typical; pterygoid fossae lengthened; auditory bullae
moderate; cheek pouches absent or very small; tail scaly, more or
less naked, cosmopolitan (_Alston_). Three molars in each jaw, the
first of which is the largest and the hinder one the least. I think
that, with the exception of the islands of the Pacific Ocean, some
of the members of this family are known in every quarter of the globe.
_GENUS MUS_.
"Muzzle pointed; eyes prominent; ears rather large, sub-naked; fur
soft (rarely mixed with spines); pollex rudimentary; claws short;
tail moderate or long, scaly, with scattered hairs; no cheek pouches;
skull elongate, narrow; temporal ridges nearly parallel; palate
compressed; incisive foramina long; auditory bullae moderately
large; coronoid process high, falcate; incisors rarely grooved;
molars with transverse ridges, each composed in youth of three
tubercles" (_Alston_).
NO. 332. MUS RATTUS.
_The Black Rat_ (_Jerdon's No. 175_).
NATIVE NAMES.--_Kala-mus_, _Kala-chuha_, Hindi; _Kala-meeyo_,
Singhalese.
HABITAT.--Chiefly Europe, but is said to be of south Asian origin;
it is stated to occur in towns near the sea-coast in India, and
Kellaart obtained it in Trincomalee only.
DESCRIPTION.--Greyish-black above, dark ashy beneath, or, as
Kellaart describes it, "above blackish-brown, along the dorsal line
nearly black; sides paler, some of the hairs with pale fulvous tips;
beneath and inside of limbs fur very short, of a uniform sooty ash
colour, separated from the colour above by a distinct line of
demarcation; ears large, rounded, slightly fulvous externally"
('Prodromus Faunae Zeylanicae,' p. 58).
[Illustration: Dentition of Black Rat.]
SIZE.--Head and body about 6-1/2 to 7-1/2 inches; tail, 7-1/2 to 8
inches.
Jerdon says of this rat that the muzzle is sharper than that of the
brown rat; the ears are more oval; it is lighter in its make, and
has much longer hair.
Whether this rat be, as Jerdon seems to suspect, imported into India
in ships or not, it is generally supposed to have had its origin in
southern Asia, and is almost identical with the Egyptian rat (_M.
Alexandrinus_). It was the common rat of England, and indeed of
northern Europe, whence it was expelled by its formidable rival, the
brown rat, before which it has gradually receded, and it is seldom
found now in England.
NO. 333. MUS DECUMANUS.
_The Brown Rat_ (_Jerdon's No. 176_).
NATIVE NAMES.--_Ghur-ka-chuha_, Hindi; _Demsa-indur_, Bengali;
_Manei-ilei_, Canarese; _Gaval-meeyo_, Singhalese.
HABITAT.--Throughout India, Ceylon, and in some parts of Burmah.
DESCRIPTION.--Fur greyish-brown, mixed with tawny above, with
longer piles of a dark colour, almost black; ears round; tail
generally longer than head and body, scaly, with short bristles at
the margins of the rings.
SIZE.--Head and body, from 8 to 10 inches; tail, from 6 to 11 inches.
The brown rat of India is identical with that of Europe, most
naturalists being now agreed that it originally came from the East.
It was supposed by Pallas that the brown rat crossed over into Russia
about the year 1727. When frightened by an earthquake, numbers swam
over the Volga from countries bordering on the Caspian Sea. It seems
to have driven out the black rat before it wherever it made its
appearance. In England it was introduced by shipping about the middle
of the last century, and has since then increased to such an extent
as to swarm over the whole country, and render the old English black
rat a comparatively rare animal. From its ferocity and fecundity the
brown rat is a veritable pest; if it cannot beat a retreat from an
enemy it will show most determined fight, and in large numbers will
attack and kill even men. A story is related by Robert Stephenson,
the great engineer, that in a coal-pit in which many horses were
employed, the rats, allured by the grain, had gathered in large
numbers. On the pit being closed for a short time, and the horses
being brought up, the first man who descended on the re-opening of
the work was killed, and devoured by the starving rats. Similar
stories have been told of men in the sewers of Paris. In the horse
slaughterhouses at Montfaucon in Paris, the rats swarm in such
incredible numbers that the carcases of horses killed during the day
would be picked clean to the bone during the night; sometimes upwards
of thirty horses would be so devoured. This shows the carnivorous
tendencies of these abominable pests. I confess to a general love
for all animals, but I draw the line at rats. There is something
repulsive about one of these creatures, and a wicked look about his
large protruding eye, like a black glistening bead, and his ways are
not pleasant; instead of keeping, as he ought, to sweet grain and
pleasant roots, he grubs about for all the carrion and animal matter
he can get.
I find there is no bait so enticing to the brown rat as a piece of
chicken or meat of any kind. I have heard stories of their attacking
children, and even grown-up people when asleep, but I cannot vouch
for the truth of this beyond what once happened to myself. I was then
inhabiting a house which swarmed with these creatures, and one night
I awoke with a sharp pain in my right arm. Jumping up, I disturbed
a rat, who sprang off the bed, and was chased and killed by me. I
found he had given me a nip just below the elbow. I once had a most
amusing rat-hunt in the house I now occupy. I had then just taken
it over on the part of the Government, in 1868. The whole building
is floored with polished marble, which, being new, was like
looking-glass. I found an enormous rat, which I took for a bandicoot,
in one of the bath-rooms, and, shutting him in for a while, I closed
the doors of a very large room adjoining, which was quite empty, and
then turned my friend in with a small black-and-tan terrier. The
scrimmage that ensued was most laughable, as both rat and dog kept
slipping and sliding all over the place. At last the former was pinned
in a corner, where he made a most determined stand, and left several
marks before he died. They seldom now come so high as the third story,
but we had two or three last year which dug a hole through a brick
wall into my study, and they were surreptitiously disposed of unknown
to my eldest little girl, whose passionate love for every living
creature made her take even the rats under her protection, and one
of them would come out every morning in the verandah to be fed by
her with crumbs and grain. This one was spared for a while, but I
was not sorry to find one day that it had fallen into a tub of water
in a bath-room and was drowned.
The brown rat breeds several times in the year, and has from ten to
fourteen at a time, and it is to be hoped that there is considerable
mortality amongst the infants. I have never kept rats as pets, but
have noticed amongst mice a tendency on the part of the mother to
devour her offspring. I have no doubt that this also is the case with
the brown rat, and aids in keeping down its numbers. It is stated
that they will attack, kill, and eat each other. The Rev. J. G. Wood
remarks in his Natural History: "From some strange cause the male
rats far outnumber the females, the proportion being about eight of
the former to three or four of the latter. This disproportion of the
sexes may possibly be caused by the cannibalistic habits of the rat,
the flesh of the female being more tender than that of the opposite
sex. Whatever may be the cause, it is clear that the wider increase
of these creatures is greatly checked by the comparative paucity of
females." During the late siege of Paris by the Germans, amongst the
various articles of food which necessity brought into use, rats held
a high place as a delicacy. It is a difficult matter to stop the
burrowing of rats; the best plan is to fill the holes with Portland
cement mixed with bits of bottle glass broken in small pieces. It
is said that quicklime will temporarily prevent rats from entering
a hole, as the lime burns their feet. A friend of mine lately told
me of some wonderful Japanese bird-lime which he uses. It is spread
on a board, and will retain any rat that puts even one foot on it.
An albino variety is common, and is sold for pets. Rats are partial
to certain scents, and some are consequently used by trappers. In
Cooley's 'Cyclopaedia' the following receipts are given:--
1. Powdered cantharides steeped in French brandy. It is said that
rats are so fond of this that if a little be rubbed on the hands they
may be handled with impunity.
2. Powdered assafoetida 8 grains, oil of rhodium 2 drams, oil of
aniseed 1 dram, oil of lavender 1/2 dram. Mix by agitation.
3. Oil of aniseed 1/2 ounce, tincture assafoetida 1/4 ounce.
4. Oil of aniseed 1/4 ounce, nitrous acid 2 to 3 drops, musk
(triturated with a little sugar) 1 grain.
These scents are not only rubbed on traps, but a few drops are mixed
with the various rat poisons, of which perhaps the most efficacious
is phosphorous paste.
NO. 334. MUS ANDAMANENSIS.
_The Andaman Rat_.
HABITAT.--The Andaman and Nicobar islands.
DESCRIPTION.--A little darker on the back than _Mus decumanus_,
paler on the sides, and dull white below. "The long piles are at once
distinguished by their flattened spinous character, which is also
slightly the case in _M. rattus_, though much less conspicuously than
in the present species. It would appear to be a burrower in the
ground" (_Blyth_). Ears round as in the brown rat.
SIZE.--Head and body, about 8 inches; tail the same.
NO. 335. MUS ROBUSTULUS.
_The Burmese Common Rat_.
HABITAT.--British Burmah.
DESCRIPTION.--Dark-brown above, under-parts whitish, stoutly
formed, with tail not quite so long as head and body; feet
conspicuously white.
SIZE.--Head and body, about 6 inches; tail, a little shorter.
Mr. Mason remarks of this rat that they are only second to the white
ants for the mischief they perpetrate. "They burrow in the gardens,
and destroy the sweet potatoes; they make their nests in the roofs
by day, and visit our houses and larders by night. They will eat into
teak drawers, boxes, and book-cases, and can go up and down anything
but glass. In the province of Tonghoo they sometimes appear in
immense numbers before harvest, and devour the paddy like locusts.
In both 1857 and 1858 the Karens on the mountains west of the city
lost all their crops from this pest." They seem to migrate in swarms,
and cross rivers by swimming. Mr. Cross captured one out of a pair
he observed swimming the Tenasserim river at a place where it is more
than a quarter of a mile wide. _M. Berdmorei_ is the same as this
species.
* * * * *
The following three are Burmese rats collected by Dr. Anderson during
the Yunnan Expedition, and are new species named by him:--
NO. 336. MUS SLADENI.
_Sladen's Rat_.
HABITAT.--Kakhyen hills; Ponsee at 3500 feet.
DESCRIPTION.--Head rather elongated; snout somewhat elongate;
muzzle rather deep; ears large and rounded, sparsely clad with short
hairs; feet well developed, hinder ones rather strong; claws
moderately long and sharp; the feet pads markedly developed,
indicating an arboreal habit of life; tail slightly exceeding length
of head and body, coarsely ringed, there being three rings to each
one-tenth of an inch; the hairs sparse and brown; general colour of
upper surface reddish-brown, more rufous than brownish, palest on
the head, many hairs with broad yellow tips; cheeks greyish-rufous;
chin, throat, and chest whitish, also the remaining under-parts, but
with a tinge of yellowish; ears and tail pale brownish. (Abridged
from Anderson's 'Anat. and Zool. Res.' p. 305.)
SIZE.--Head and body of one, about 6.30 inches; tail, 7.20 inches.
Dr. Anderson says this species is closely allied to Hodgson's _Mus
nitidus_, but its skull is less elongated, with a shorter facial
portion, with very much shorter nasals, and with a more abruptly
defined frontal contraction than either in _M. nitidus_ or _M.
rufescens_ so called. He adds that this appears to be both a tree
and a house rat.
NO. 337. MUS RUBRICOSA.
_The Small Red Rat of the Kakhyen Hills_.
HABITAT.--Kakhyen hills and the Burma-Chinese frontier at Ponsee,
and in the houses of the Shan Chinese at Hotha.
DESCRIPTION.--"Snout moderately pointed and long; ears small, and
somewhat pointed; hind foot long and narrow; claws moderately long,
compressed and sharply pointed; upper surface dark rusty brown,
darkest on the middle and back, and palest on the muzzle, head and
shoulder; on the sides and lower part of shoulder the reddish brown
tends to pass into greyish; feet greyish; the sides of the snout
greyish; all the under-parts silvery grey tending to white, without
any trace of rufous, or but with a very faint yellowish blush; the
tail, dull brown, is somewhat shorter than the body and head, and
it is coarsely ringed, 2-1/2 rings to one-tenth of an inch, the hair
being short, sparse, and dark brown" ('Anat. and Zool. Res.' p. 306).
SIZE.--Head and body, 5.70 inches; tail, 5.15 inches.
NO. 338. MUS YUNNANENSIS.
_The Common House Rat of Yunnan_.
HABITAT.--Yunnan, at Ponsee; Hotha and Teng-yue-chow.
DESCRIPTION.--"Muzzle rather short and broad; ear large and rounded,
its height considerably exceeding the distance between the inner
canthus and the front of the muzzle, sparsely clad with short hairs;
feet well developed; hind foot moderately long; pads prominent;
claws compressed, strong, curved, and sharp; tail coarsely ringed,
three rings to one-tenth of an inch; upper surface dark rich brown,
with intermixed pale hairs, with broad brown tips, the sides of the
face below the moustachial area, chin, throat, and all the
under-parts yellowish washed with rufous; the ears and tail dusky
brown; feet pale yellowish, and more or less brownish above; the tail
varies in length, but is generally longer than the body and head,
although it may occasionally fall short of that length" ('Anat. and
Zool. Res.' pp, 306, 307).
SIZE.--Head and body, 5.70 inches; tail, 5.65 inches. An adult female
had a much longer tail.
NO. 339. MUS INFRALINEATUS.
_The Striped-bellied Rat_ (_Jerdon's No. 178_).
HABITAT.--Madras; Bustar forests.
DESCRIPTION.--"Above, the fur fulvous, with the shorter hairs lead
coloured; throat, breast, and belly pure white, with a central pale
fulvous brown streak; tail slightly hairy."--_Jerdon_.
SIZE.--Head and body, 5-1/2 inches; tail, not quite 5 inches; another
about 5 inches; tail, 4-1/4 inches.
Jerdon calls this a field rat in his popular name for it, but I think
that the term should be restricted to the _Nesokia_ or true field
and earth-burrowing rats. He is of opinion that Gray's _Mus
fulvescens_ from Nepal is the same, the description tallying to some
extent, concluding with: "in one specimen a central yellow streak,"
i.e. on the belly.
NO. 340. MUS BRUNNEUS.
_The Tree Rat_ (_Jerdon's. No. 179_).
HABITAT.--India and Ceylon. The common house rat of Nepal.
DESCRIPTION.--Above rusty brown; below rusty, more or less
albescent; extremities pale, almost flesh-coloured; ears rather
long; head rather elongated; tail equal to and sometimes exceeding
head and body.
SIZE.--Head and body, from 8-1/2 to 9-1/2 inches; tail, from 9 to
9-1/2 inches.
Jerdon states that this rat, which Dr. Gray considered identical with
_M. decumanus_ (_see_ 'Ann. and Mag. Nat. Hist.' vol. xv. 1845, p.
267), "is to be found throughout India, not habitually living in
holes, but coming into houses at night; and, as Blyth remarks, often
found resting during the day on the _jhil-mil_ or venetian blinds.
It makes a nest in mango-trees or in thick bushes and hedges. Hodgson
calls it the common house rat of Nepal, and Kellaart also calls it
the small house rat of Trincomalee." It is probable that this is the
rat which used to trouble me much on the outskirts of the station
of Nagpore. It used to come in at night, evidently from outside, for
the house was not one in which even a mouse could have got shelter,
with masonry roof, and floors paved with stone flags. Kellaart
evidently considered it as distinct from _M. decumanus_, which he
stated to be rare in houses in the town of Trincomalee, though
abundant in the dockyard.
NO. 341. MUS RUFESCENS.
_The Rufescent Tree Rat_ (_Jerdon's No. 180_).
NATIVE NAMES.--_Gachua-indur_, Bengali; _Ghas-meeyo_, Singhalese.
HABITAT.--India generally; Ceylon.
DESCRIPTION.--Fur above pale yellowish-brown; under fur lead
coloured, mixed with longer piles of stiff, broad, plumbeous black
tipped hairs; head long; muzzle narrow; whiskers long and black; ears
large, subovate, slightly clad with fine hairs; eyes large; incisor
teeth yellow; feet brownish above, but the sides and toes are
whitish; tail longer than head and body.
SIZE.--Head and body, from 5-1/2 to 7-1/2 inches; tail from 6-1/2
to 8-1/2 inches.
This is _M. flavescens_ of Elliot, and is so noticed in Kellaart's
'Prodromus.' He calls it "the white-bellied tree-rat of Ceylon," and
he states that it lives on trees or in the ceiling of houses in
preference to the lower parts. Sir Walter Elliot observed it chiefly
in stables and out-houses at Dharwar. According to Buchanan-Hamilton
it makes its nests in cocoanut-trees and bamboos, bringing forth five
or six young in August and September. "They eat grains, which they
collect in their nests, also young cocoanuts. They enter houses at
night, but do not live there." Kellaart's _M. tetragonurus_ is a
variety of this, if not identical.
NO. 342. MUS NIVEIVENTER.
_The White-bellied House Rat_ (_Jerdon's No. 181_).
HABITAT.--The lower Himalayan ranges.
DESCRIPTION.--"Above blackish-brown, shaded with rufous; below
entirely pure white, tail and all."--_Blyth_.
SIZE.--Head and body, 5-1/4 to 7 inches; tail, 6 to 7-1/2 inches.
Hodgson stated this to be a house rat in Nepal, but not very common.
Jerdon found it common at Darjeeling. Specimens have been received
from Mussoorie.
NO. 343. MUS NITIDUS.
_The Shining Brown Rat_ (_Jerdon's No. 182_).
HABITAT.--Nepal; Darjeeling.
DESCRIPTION.--Dusky brown above, dusky hoary below. According to
Hodgson it is "distinguished for its smooth coat or pelage, wherein
the long hairy piles are almost wanting. It is a house rat, like _M.
niveiventer_, but much rarer, and frequents the mountains rather
than the valleys." The long hairs are 11/16 inch in length, horny
at the base, with black tip, the short fur ashy, with rufous tips.
SIZE.--Head and body, 6-1/2 inches; tail 7-1/4 inches.
Blyth writes of this species ('J. A. S. B.' vol. xxxii. 1863, p. 343):
"We have several specimens of what I take to be this rat from
Darjeeling. They are especially distinguished by the fineness and
softness of the fur. One specimen only, of eight from Darjeeling,
which I refer to this species, has the lower parts pure white,
abruptly defined."
There is a smaller rat, only four inches in length, which agrees
exactly with the above, which Hodgson named _M. horietes_. It is not
mentioned in Blyth's Catalogue, but it has not been overlooked by
Blyth, as Jerdon's remarks would lead one to suppose, for in the
'Memoir on the Rats and Mice in India,' by the former, in the 'J.
A. S. B.' vol. xxxii. for 1863, it is entered with a quotation from
Hodgson.
NO. 344. MUS CAUDATIOR.
_The Chestnut Rat_ (_Jerdon's No. 183_).
HABITAT.--The lower Eastern Himalayas, i.e., Nepal, Darjeeling,
&c.; also in Burmah, Lower Pegu, and Martaban.
DESCRIPTION.--"Above a fine bright cinnamon colour, with
inconspicuous black tips; the under-parts white, which is abruptly
divided from the cinnamon hue above" (_Blyth_). Sometimes
yellowish-white (_Jerdon_). Muzzle sharp; ears and tail long.
SIZE.--Head and body, about six inches; tail, 7-3/4 inches.
According to Blyth the Nepal specimens are darker than those from
Burmah, which he says "differs only from the Nepalese animal of Mr.
Hodgson by having the upper parts entirely of a bright cinnamon
colour."
NO. 345. MUS CONCOLOR.
_The Common Thatch Rat of Pegu_.
HABITAT.--Upper and Lower Burmah, Malayan peninsula.
DESCRIPTION.--I have been unable to trace any accurate description
of this rat, which Blyth says "conducts from the long-tailed arboreal
rats to the ordinary house mice." In his 'Catalogue of the Mammals
of Burmah,' published in the 'Jour. Asiatic Soc. Beng.' for 1875,
he remarks that "it requires to be critically examined in the fresh
state." In the 'J. A. S. B.,' vol. xxviii. p. 295, he describes a
young one as dark greyish mouse colour; but this is not reliable,
as the young rats and mice change colour as they attain full
growth.[23]
[Footnote 23: Since writing the above, Dr. Anderson has kindly
allowed me to examine the specimens of _Mus concolor_ in the museum,
and in the adult state they are considerably more rufescent. In one
specimen, allowing for the effects of the spirit, the fur was a bright
rufescent brown; but, whatever be the tint of the prevailing colour,
it pervades the whole body, being but slightly paler on the
under-parts. Size, about 4 inches; tail, about 4-1/2 inches.--R. A.
S.]
NO. 346. MUS PALMARUM.
_The Nicobar Tree Rat_.
HABITAT.--Nicobar Islands.
NO. 347. MUS CEYLONUS.
HABITAT.--Ceylon.
DESCRIPTION.--Fur soft, lead colour; hair of upper parts tipped with
dark fawn and black; ears large, naked; whiskers long, black; tail
longer than the head and body, scaly.
SIZE.--Head and body, 4-3/4 inches; tail, 6 inches.
"This small rat is found in out-houses in the cinnamon gardens at
Colombo. I have no reason to think it to be the young of the former
species (_M. decumanus_); the teeth were well developed; the darker
colour and long tail will easily distinguish the species from other
Colombo rats" (_Kellaart_). The character of the molar teeth is all
that can be depended on in the foregoing description, and this may
require further investigation. The young of rats and mice are always
darker than the adults, and the tail is longer in proportion.
* * * * *
The following are doubtful species:--
NO. 348. MUS PLURIMAMMIS.
_Jerdon's No. 177_.
This, which Blyth considered a good species, is, I am informed,
referable with _M. Taraiyensis_ and _M. Morungensis_ to Gray's
_Nesokia Bengalensis_. The type and drawing of it are in the British
Museum.
NO. 349. MUS AEQUICAUDALIS.
of Hodgson, described in Horsfield's Catalogue as pure dark brown
above, with a very slight cast of rufescent in a certain aspect;
underneath from the chin to the vent, with interior of thighs,
yellowish-white; ears nearly an inch long; head proportionately long
('Ann. and Mag. Nat. Hist.' new series, iii. p. 203). This, with
Blyth's _M. nemoralis_, seems identical with _M. brunneus_.
_Mus arboreus_ of Horsfield's Catalogue is _Mus rufescens_. It
remains to be seen whether there is sufficient difference between
_M. rufescens_ and _M. niveiventer_ to warrant the separation of the
latter as a distinct species.
* * * * *
The following species lead on to the mice--beginning with the
long-tailed arboreal species, _Vandeleuria_ of Gray, which connect
the arboreal rats with the house mice.
The characteristics of _Vandeleuria_ are: upper incisors triangular,
grooved in front; ears hairy; fur soft, with long bristles
interspersed; long tail, sparsely haired; hind feet very long,
slender; soles bald beneath; toes .45 long, slender, compressed, the
pads much more strongly developed than in ground mice; the inner and
outer toes with a small flattened nail.
NO. 350. MUS OLERACEUS.
_The Long-tailed Tree Mouse_ (_Jerdon's No. 184_).
NATIVE NAMES.--_Marad-ilei_, Canarese; _Meina-yelka_, Telegu of the
Yanadees (_Jerdon_).
HABITAT.--Throughout India from north to south, but has not been
reported from Ceylon. In Burmah Dr. Anderson found it in the valley
of the Nampoung, a frontier stream dividing Burmah from China.
DESCRIPTION.--Upper surface rich rufous or chestnut red, paling to
brown on the ears and muzzle before the eyes; under-parts white, with
a yellowish tinge; feet pale brown, shading off into white on the
toes; under surface of feet yellowish; tail brownish or dusky with
grey hairs; it tapers to a point, finely ringed; sparsely haired
between the rings, the hairs more numerous and longer towards the
tips. The length of the head, according to Dr. Anderson, whose
description ('Anat. and Zool. Res.' p. 313) is more complete than
Jerdon's, is about one-third the length of the body; the muzzle is
moderately long and slightly contracted behind the moustachial area;
eyes large; ears ovate, sparsely clad.
SIZE.--Head and body, from 2-1/2 to 3 inches; tail one-half longer
than the combined length of body and head.
Jerdon says of this pretty little mouse that "it is most abundant
in the south of India, where it frequents trees, and very commonly
palm-trees, on which it is said to make its nest generally. It,
however, occasionally places its nest in the thatch of houses, on
beams, &c. It is very active, and from its habits difficult to
procure" ('Mammals of India,' p. 202). According to Sykes it
constructs its nest of oleraceous herbs in the fields, and Hodgson
states it to tenant woods and coppices in Nepal.
NO. 351. MUS NILAGIRICUS.
_The Neilgherry Tree Mouse_ (_Jerdon's No. 185_).
HABITAT.--Ootacamund.
DESCRIPTION.--"Above deep but bright chestnut brown, beneath bright
fawn yellow, with a distinct line of demarcation between the two
colours; head rather elongated; ears long, oval; tail somewhat
hairy."--_Jerdon_.
SIZE.--Head and body, 3-1/2 inches; tail, 5 inches.
This tree mouse was discovered and named by Dr. Jerdon. He says: "The
first I observed was brought into the house by a cat. I afterwards,
on two or three occasions, found the nest, a mass of leaves and grass,
on shrubs and low trees, from four to five feet from the ground, and
on one occasion it was occupied by at least eight or ten apparently
full-grown mice."
NO. 352. MUS BADIUS.
_The Bay Tree Mouse_.
HABITAT.--The valley of the Sittang, Burmah.
DESCRIPTION.--"Similar to _M. oleraceus_, but with the eye fully
twice as large, and black whiskers; colour of the upper parts a more
rufous chestnut or cinnamon hue, of the lower parts white, almost
pure."--_Blyth_.
SIZE.--Head and body, 3 inches; tail, 4-3/8 inches.
NO. 353. MUS GLIROIDES.
_The Cherrapoonjee Tree Mouse_.
HABITAT.--Khasia hills.
DESCRIPTION.--Fur exceedingly dense and fine, of a light brown,
tinged with fawn; the basal two-thirds of the piles are dusky ash
coloured; the lower parts are white, very faintly tinged with fawn;
the white purest about the lips and chin; whiskers long; feet large
and sparsely clad with white hairs; a distinct brown mark on each
hind foot reaching almost to the division of the toes; ears smallish,
ovoid, naked.
SIZE.--Head and body, 2 inches; tail (?) mutilated.
Blyth says this animal has much of the aspect of the European dormouse
(_Myoxus avellanarius_), but nothing is said about its dentition,
which would at once settle the question whether the young specimen
with its imperfect tail were a true _Mus_ or a species of
_Myoxus_.[24]
[Footnote 24: See Appendix A for description and dentition of
_Myoxus_.]
NO. 354. MUS PEGUENSIS.
_The Pegu Tree Mouse_.
HABITAT.--The Sittang valley, Burmah.
DESCRIPTION.--Fulvescent olive brown on the upper parts,
yellowish-white below; whiskers remarkably long; the tail very long
and conspicuously haired towards the tip; more so, Blyth remarks,
than any other mouse, especially when held up to the light.
SIZE.--Head and body, 3-1/8 inches; tail, 3-7/8; in one specimen,
4-1/2 inches.
* * * * *
We now come to the terrestrial or house mice.
NO. 355. MUS URBANUS.
_The Common Indian Mouse_ (_Jerdon's No. 186_).
NATIVE NAMES.--_Lengtia-indur_, Bengali; _Mesuri_, _Musi_, _Chuhi_,
Hindi.
HABITAT.--Throughout India and Ceylon.
DESCRIPTION.--Somewhat resembling the English mouse, but with very
much longer, coarser tail, larger eyes, and smaller ears; dusky
reddish-brown above, somewhat paler below; the feet paler still,
whitish in some; the tail nude, thick at base, longer by an inch than
the head and body, and of a dark brown colour. The young are more
dusky.
SIZE.--Head and body, about 2 to 3 inches; tail, 3 to 4 inches.
I have kept these mice in confinement for considerable periods, and
have had many opportunities of studying their habits of late. During
many years' residence in the Currency Office, I never once found a
mouse in my private quarters on the third story, although I
frequently observed them in the vaults and strong rooms on the ground
floor. During my absence at Simla in 1880 my quarters were unoccupied,
as the Public Works Department were giving the building a thorough
repair. It was then, I suppose, a few of the mice from the ground
floor were driven upstairs, and, being unmolested by us, as we liked
to see the little things playing about, they increased to a most
uncomfortable extent within eight months. I failed to discover their
breeding places, though I suspect they made much use of a large
doll's-house for the purpose, for on taking out the front staircase,
under which the bells of the establishment were hung, I found a nest
of torn paper, and I caught two young ones in one of the rooms. Some
of them came out every night whilst we were at dinner, and paid a
visit to a rose-headed parraquet (_Palaeornis rosa_), mounting up
on Polly's perch, and sitting down to supper in the tin receptacles
for food at each end. She generally treated them with silent contempt,
or gave a snappish little peck if they were too familiar; sometimes,
when they were too sky-larky, she retreated to her ring above, where
she swung and looked down at them from a coign of vantage. Their
agility in running up and down the wires of a cage is marvellous.
They have also an extraordinary faculty for running up a
perpendicular board, and the height from which they can jump is
astounding. One day, in my study, I chased one of these mice on to
the top of a book-case. Standing on some steps, I was about to put
my hand over him, when he jumped on to the marble floor and ran off.
I measured the height, and have since measured it again, 8 feet 9-1/2
inches.
I consider this species the most muscular of all mice of the same
size. I have had at the same time in confinement an English mouse
(albino), a Bengal field mouse, and house mice from Simla of another
species, and none of them could show equal activity. I use, for the
purpose of taming mice, a glass fish-globe, out of which none of the
other mice could get, but I have repeatedly seen specimens of _M.
urbanus_ jump clear out of the opening at the top. They would look
up, gather their hind quarters together, and then go in for a high
leap. They are much more voracious than the Simla or other mice. The
allowance of food given would be devoured in less than half the time
taken by the others, and they are more given to gnawing. What sort
of mothers they are in freedom I know not, but one which produced
four young ones in one of my cages devoured her offspring before they
were a week old. I have two before me just now as I write, and they
have had a quarrel about the highest place on a little grated window.
The larger one got the advantage, so the other seized hold of her
tail, and gave it a good nip.
* * * * *
Now we come to some doubtful species, doubtful in the sense that they
should not be separated, but considered as one to be named afterwards,
according to priority of discovery. Dr. Anderson is at present
investigating the matter, and we must await his decision, but from
such external observations as I have been able to make, it appears
probable that the following will prove identical:--
_Mus homourus_; _Mus Darjeelingensis_; _Mus Tytleri_; _Mus
Bactrianus_; _Mus cervicolor_(?)--_Jerdon's Nos. 187, 189, 190, 191,
and 192_. These are all hill mice, except the last, and found under
the same conditions.
NO. 356. MUS HOMOURUS.
HABITAT.--Lower Himalayan range.
DESCRIPTION.--Dark rufescent above, rufescent white below; hands
and feet fleshy white; tail equal to length of head and body; "fur
more gerbille-like in character than in _M. musculus_" (or
_urbanus_), stated to be the common house mouse of the Himalayan hill
stations from the Punjab to Darjeeling. Stated by Hodgson to have
eight teats only in the female, other mice having ten. Possibly his
description was founded on young specimens. I myself was of opinion
for some time that I had got two species of hill mice, a larger and
a smaller, the latter being so much darker in colour, but I kept them
till the young ones attained full size in six months, at which time
they were not distinguishable from the old ones. Hodgson may have
overlooked the pectoral mammae when he noted the number.
SIZE.--Head and body, 3-1/2 inches; tail, 3-1/2 inches.
NO. 357. MUS DARJEELINGENSIS.
DESCRIPTION.--Dusky brown, with a slight chestnut reflection;
under-parts pale yellowish-white.
SIZE.--Head and body, 3 inches; tail, 2-1/2 inches.
NO. 358. MUS TYTLERI.
HABITAT.--Dehra Doon.
DESCRIPTION.--Fur long and full, pale, sandy mouse-coloured above,
isabelline below; pale on the well-clad limbs, and also on the tail
laterally and underneath.
SIZE.--Head and body, 2-3/4 inches; tail, 2-3/4 inches.
NO. 359. MUS BACTRIANUS.
HABITAT.--Punjab, Kashmir, Candahar, Baluchistan, and Southern
Persia.
DESCRIPTION.--Upper parts brown above, with a sandy tinge, more on
the head; the longer hairs with a dusky tip; the basal two-thirds
deep ash; under-parts and feet white; tail clad thinly with fine
whitish hair; the fur in general long, dense, and silky.
SIZE.--Head and body, from 2-1/4 to 3-1/4 inches; tail, about the
same.
This is the mouse, I think, that I caught in the house at Simla in
1880. Of eight specimens I got--seven in a cupboard in the
dining-room and one in a bath-room--I sent two in spirits to the
Indian Museum and brought down to Calcutta three alive, which I kept
for about seven months, when they died. I have since then seen living
specimens of _M. bactrianus_ from Kohat, with which they appear to
be identical. They also resemble--I speak under correction--_M.
cervicolor_, which is a field mouse found in Bengal. I made the
following notes regarding them: Fur very fine, close and silky,
rufescent brown, more rufous on the head, isabelline below; feet
flesh-coloured, hinder ones large, much larger than those of the
English mouse; the hind-quarters are also more powerful; has a very
pretty way of sitting up, with the body bent forwards, and its hands
clasped in an attitude of supplication. The young mice seem darker
both above and below, and are much more shy than the old ones, of
which one soon after being caught took bits of cake from my fingers
through the bars of its cage. More delicate looking than _Mus
urbanus_, with a much shorter and finer tail; less offensive in
smell.
Dr. Anderson got, not long ago, two of these mice in a box from Kohat.
They bore the journey uncommonly well, and were in lively condition
when I saw them at the Museum. Whilst we were talking about them,
we noticed an act of intelligence for which I should not have given
them credit had I not seen it with my own eyes. They were in a box
with a glass front; in the upper left-hand corner was a small sleeping
chamber, led up to by a sloping piece of wood. The entrance of this
chamber was barred by wires bent into the form of a lady's hair-pin,
and passed through holes in the roof of the box.
The mice had been driven out, and the sleeping-chamber barred, for
they were having their portraits taken. Whilst we were talking we
found, to our surprise, that one mouse was inside the chamber,
although the bars were down. There seemed hardly space for it to
squeeze through; however, it was driven out, and we went on with our
conversation, but found, on looking at the cage again, that our
little friend was once more inside, so he was driven out again, and
we kept an eye on him. To our great surprise and amusement we saw
him trot up his sloping board, put his little head on one side, and
seize one of the wires, which worked very loosely in its socket, give
it a hitch up, when he adroitly caught it lower down, hitched it up
again and again till he got it high enough to allow him to slip in
underneath, and then he was quite happy once more. He had only been
in the box two days, so he was not long in finding out the weak point.
I begin to believe now in rats dipping their tails into oil-bottles,
and other wonderful stories of murine sagacity that one reads of.
Mice, are supposed to live from two-and-a-half to three years. I had
the English albino above mentioned for three.
NO. 360. MUS CRASSIPES.
_The Large-footed Mouse_ (_Jerdon's No. 188_).
HABITAT.--Mussoorie and, according to Jerdon, the Neilgherries.
DESCRIPTION.--This is stated to be like _M. homourus_, but the
difference is well marked in a very much longer tail and much larger
feet.
SIZE.--Head and body, 2-3/4 inches; tail, 3/4 inch; hind foot, 3/4
inch.
NO. 361. MUS SUBLIMIS.
HABITAT.--Ladakh, 13,000 feet.
DESCRIPTION.--Brown above; whitish below; the colours gradually
blending; fur soft and long; all except the tips dark slaty grey,
the terminal portions of the shorter hairs being light brown, and
of the longer hairs dark brown; upper whiskers black; lower white;
ears oval; feet thinly clad with short light brown hairs; tail with
short bristly hairs, dusky brown above, whitish below; tail longer
than head and body.
SIZE.--Head and body, 2.6 inches; tail, 3.05; length of hind foot,
0.83 inch.
Mr. Blanford, who named the above species, which was procured in the
expedition to Yarkand, is doubtful whether it may not be referable
to the last species.
NO. 362. MUS PACHYCERCUS.
HABITAT.--Yarkand.
DESCRIPTION.--Sandy brown above; under-parts white; fur soft and
very like _M. bactrianus_; ears large, rounded, hairy; feet clad
above with white hair; soles naked; tail thick, shorter than head
and body, and thinly clad with white bristles throughout; skin dark
above, pale below; incisors deep yellow.
SIZE.--Head and body, 2.35 inches; tail, 1.9 to 2 inches.
Mr. Blanford says this is a house mouse. It is figured in Blanford's
'Mammalia of the Second Yarkand Mission.'
NO. 363. MUS ERYTHRONOTUS.
HABITAT.--Yarkand, Persia.
DESCRIPTION.--Rufous, washed with blackish above, white below,
abruptly separated; hairs on the back are slaty at the base, then
blackish and bright ferruginous at the tips, the extreme points being
black, except on the sides, where the black tip is wanting; upper
whiskers black, lower white; ears large, rounded, naked; feet white
above, dusky and naked below; tail equal to head and body, nearly
naked. Mammae six.
SIZE.--Head and body, 4 inches; tail, 4.2 inches.
This mouse is figured and carefully described in Blanford's 'Eastern
Persia,' vol. ii. p. 35.
NO. 364. MUS CERVICOLOR.
_The Fawn-coloured Field Mouse_.
HABITAT.--Bengal, Nepal, Southern India.
DESCRIPTION.--"Distinguished by its short tail. Above dull fawn,
below sordid white; lining of ears and extremities pale" (_Blyth_).
"Ears large, hairy" (_Jerdon_). Of the specimens I have seen the fur
is soft and of a light sandy brown above and white below, very like
_M. bactrianus_.
SIZE.--Head and body, 3-1/2 inches; tail, 2-7/8 inches.
NO. 365. MUS TERRICOLOR.
_The Earth-coloured Field Mouse_.
HABITAT.--India generally, I think. It has been found in the valley
of the Ganges, in Bengal, in the Santal district west of Midnapore,
and Southern India.
DESCRIPTION.--The colour varies according to the soil, but in
general fawn brown, more or less rufescent--those from the valley
of the Ganges being darker than those from the ferruginous soil of
other parts. The under-parts are white, abruptly separated from the
brown; fur short and soft.
SIZE.--Head and body, 2-1/2 inches; tail, 2-1/8 inches.
NO. 366. MUS PEGUENSIS.
_The Pegu Field Mouse_.
HABITAT.--The valley of the Sitang River, Burmah.
DESCRIPTION.--"Fur very full and dense, pale fulvescent olive brown
on the upper parts, slightly yellowish-white below; whiskers
remarkably long" (_Blyth_). Tail longer than head and body, and well
clad with hairs, especially towards the tip.
SIZE.--Head and body, 3-1/8 inches; tail, nearly 4 inches.
NO. 367. MUS NITIDULUS.
_The Shiny Little House Mouse of Pegu_.
HABITAT.--The Sitang valley in Burmah.
DESCRIPTION.--The description given of this mouse by Blyth is
extremely vague. He says: "A house mouse apparently, with tail equal
to head and body, and uniformly furnished with minute setae to the
end; ears large and ample; colour nearly that of _M. decumanus_, with
the under-parts subdued white, tolerably well defined."
He remarks further on that the front teeth are conspicuously larger
than those of _M. musculus_ and _M. urbanus_.
SIZE.--Head and body, 3-1/4 inches; tail, the same.
NO. 368. MUS BEAVENI.
_Beaven's Mouse_.
HABITAT.--Maubhum, and, according to Blyth's Catalogue, Burmah,
valley of the Salween.
DESCRIPTION.--"Above rusty brown, medially black; lips and the whole
under side pale ochraceous; feet white, all the hair being slate
coloured at the base; tail above brown, below with white hairs; upper
whiskers black, lower white. Rather smaller and more delicately
built than our common harvest mouse."--_Prof. Peters_, 'P. Z. S.'
1866, p. 559.
NO. 369. MUS CUNICULARIS.
_The Little Rabbit-Mouse_.
HABITAT.--Cherrapunji, Assam.
DESCRIPTION.--"A small field (?) mouse, remarkable for its ample
ears and tail shorter than head and body; colour of a wild rabbit
above, below white; and the feet with brownish hairs above, but with
white hairs upon the toes; tail conspicuously ringed; the setae
minute and inconspicuous."--_Blyth_.
SIZE.--Head and body, 2-1/2 inches; tail, 2-1/8 inches; ears
posteriorly half an inch.
NO. 370. MUS ERYTHROTIS.
_The Cherrapunji Red-eared Mouse_.
HABITAT.--Cherrapoonji, Assam.
DESCRIPTION.--A small mouse with very deep soft fur, very long and
silky, of a rich dark brown colour, grizzled and brightly tinged with
rufous or rufo-ferruginous towards the tail, and upon the ears
conspicuously. In such spirit specimens as I have seen the colour
was darker than in life, but the soft silkiness of the fur could be
seen to advantage as it floated in the clear liquid; the lower parts
are whitish, tinged with fawn; feet with brown hairs above; ears
small and hirsute, and the tail is also hairy.
SIZE.--Head and body, 2-1/4 inches; tail, 2-3/8 inches.
NO. 371. MUS FULVIDIVENTRIS.
HABITAT.--Ceylon, Trincomalee.
DESCRIPTION.--This is a small mouse very like _Mus cervicolor_, or
perhaps _M. terricolor_, which it more nearly approaches in size.
Kellaart in his 'Prodromus,' calls it _cervicolor_, but Blyth
afterwards separated it under the name given above, though after all
I think he was doubtful whether it ought to have been so distinguished.
The fur is long, soft, and glossy, fulvous fawn brown above, paler
below; feet dingy grey.
SIZE.--Head and body, 2-9/10 inches; tail, 2-5/10 inches.
NO. 372. MUS KAKHYENENSIS.
_The Kakhyen Mouse_.
HABITAT.--Burmo-Chinese frontier, Ponsee.
DESCRIPTION.--Differs from _Mus urbanus_ by its shorter tail, longer
hind feet, and larger ears; muzzle moderately deep, and short; ears
large and rounded; fur long, dense, and soft, reddish-brown on the
upper parts, with a dark speckled appearance due to the stronger
hairs having broad brown tips; sides of the head dusky greyish; chin
to vent and under-parts greyish-white, with a silvery sheen; feet
dusky pale brown; ears and upper surface of tail dark brown, under
surface of tail pale brown.--_Anderson_.
SIZE.--Head and body, 2.90 inches; tail, 3.36 inches.
This mouse was discovered and named by Dr. Anderson, who procured
one example at Ponsee, where it occurs, he says, on the old rice and
Indian corn clearings. The next species is also a new one discovered
and named by him.
NO. 373. MUS VICULORUM.
_The Kakhyen House Mouse_.
HABITAT.--The Burmo-Chinese frontier, Ponsee.
DESCRIPTION.--Muzzle rather sharply pointed, moderately long and
not deep; ears moderately large, rounded; its height a little in
excess of the distance between the inner canthus and the front of
the muzzle; hind-feet not long; tail a little longer than the body
and head, finely ringed, five rings to one-tenth of an inch; fur soft,
short, dense, dull dark brown on the upper parts, tending to blackish
on the back, paling to brownish on the sides, and passing into pale
dusky brownish on the under parts with a silvery sheen; feet
brownish; toes with shining greyish-yellow hairs; ears and tail
brown. (_See_ Anderson's 'Anat. and Zool. Res.,' p. 308.)
SIZE.--Head and body, 2-9/10 inches; tail, 3.14 inches.
This species, according to Dr. Anderson, frequents the villages and
houses of the Kakhyens. He obtained it at Ponsee.
* * * * *
We now come to an interesting little group of mice, of which the hairs
are mixed with flat spines, which form the genus _Leggada_ of Gray,
a term taken from the Wuddur name for the next species.
_GENUS LEGGADA_.
CHARACTERISTICS.--Molars high, with somewhat convex crowns; the
cross ridges of the upper grinders deeply three-lobed; the front one
with an additional lunate lobe at the base of its front edge; fur
fine, mixed with numerous spines somewhat flattened.
NO. 374. LEGGADA PLATYTHRIX.
_The Brown Spiny Mouse_ (_Jerdon's No. 194_).
NATIVE NAMES.--_Leggade_ and _Kal-yelka_, of Wuddurs; _Gijeli-gadu_,
Telegu, of Yanadees; _Kal-ilei_, Canarese.
HABITAT.--Southern India.
DESCRIPTION.--Sandy brown or light brown fawn above, white
underneath, with a band of pale fawn separating the two colours.
The fur mixed with flat transparent spines, smaller beneath; head
long; muzzle pointed; ears rather large, oblong, rounded, about half
an inch in length.
SIZE.--Head and body, 3-1/2 inches; tail, 2-1/2 inches.
The following description has been given by Sir Walter Elliot and
reproduced in Jerdon's 'Mammals': "The Leggade lives entirely in the
red gravelly soil in a burrow of moderate depth, generally on the
side of a bank. When the animal is inside the entrance is closed with
small pebbles, a quantity of which is collected outside, by which
its retreat may always be known. The burrow leads to a chamber in
which is collected a bed of small pebbles on which it sits, the thick
close hair of the belly protecting it from the cold and asperity of
such a seat. Its food appears to be vegetable. In its habits it is
monogamous and nocturnal.
"In one earth which I opened, and which did not seem to have been
originally constructed by the animal, I found two pairs, one of which
were adults, the other young ones about three-parts grown. The mouth
of the earth was very large, and completely blocked up with small
stones; the passage gradually widening into a large cavity, from the
roof of which some other passages appeared to proceed, but there was
only one communication with the surface, viz. the entrance. The old
pair were seated on a bed of pebbles, near which, on a higher level,
was another collection of stones probably intended for a drier
retreat; the young ones were in one of the passages, likewise
furnished with a heap of small stones."
Dr. Jerdon adds he has often opened the burrows of this mouse, and
can confirm the above account. He also states that the Yanadees of
Nellore declare that one variety uses small sticks instead of stones
to sit upon, and they give it a distinct appellation, but he could
not detect any difference in the specimens they brought him.
NO. 375. LEGGADA SPINULOSA.
_The Dusky Spiny Mouse_ (_Jerdon's No. 195_).
HABITAT.--Punjab, and also Southern India.
DESCRIPTION.--"Nearly affined to _M. platythrix_ (Sykes), but of a
dark dusky colour above, with fulvous tips to the softer fur; below
and all the feet dull whitish; upper rodential tusks orange, the
lower white; whiskers long and fine, the posterior and longer of them
black for the basal half or more, the rest white."--_Blyth_, 'J. A.
S. B.' 1863.
SIZE.--Head and body, 3-3/4 inches; tail, 3 inches.
NO. 376. LEGGADA JERDONI.
_The Himalayan Spiny Mouse_ (_Jerdon's No. 196_).
HABITAT.--Himalayan range, up to 12,000 feet.
DESCRIPTION.--"Bright dark ferruginous above, pure white below;
some fine long black tips intermingled among the spines of the back;
limbs marked with blackish externally; the feet white."--_Blyth's_
'Mem., J. A. S. B.' vol. xxxii.
SIZE.--Head and body, 4 inches; tail, 3-1/2 inches.
Dr. Jerdon first found this mouse at Darjeeling, but afterwards in
the valley of the Sutlej in Kunawur, at an elevation of nearly 12,000
feet, living under large stones.
NO. 377. LEGGADA LEPIDA.
_The Small Spiny Mouse_ (_Jerdon's No. 197_).
NATIVE NAMES.--_Chitta-burkani_, _Chit-yelka_, _Chitta-ganda_,
Telegu of Wuddurs; _Chitta-yelka_ of Yanadees.--_Jerdon_.
HABITAT.--Southern India.
DESCRIPTION.--Similar to _L. platythrix_, but smaller and more
weakly spinous; above pale sandy brown, pure white below, the two
colours clearly separated. "The spines are small, fine, transparent,
and of a dusky tinge, tipped with fawn; head very long; muzzle
pointed; ears large, ovate, naked; tail naked, limbs rather long,
fine."--_Jerdon_.
SIZE.--Head and body, 2-1/2 to 3 inches; tail, 2-3/4 inches.
Jerdon says of this mouse that he has found it in gravelly soil in
gardens and woods in most parts of Southern India making a small
burrow, which generally has a little heap of stones placed at a short
distance from the hole. It is preyed on now and then by the common
Indian roller or jay, and it is very generally used as a bait to catch
that bird with bird-lime.
_GENUS GOLUNDA_.
The following rats are separated by Gray as a distinct genus, which
from the Canarese name of the type he has called _Golunda_, the
characteristics of which are: "the grinders, when perfect, low, with
a broad, flat crown; the cross ridges of the crown of the upper
grinders divided into three distinct slightly raised tubercles;
upper incisors grooved; rest like _Mus_."
NO. 378. GOLUNDA ELLIOTI.
_The Bush Rat or Coffee Rat_ (_Jerdon's No. 199_).
NATIVE NAMES.--_Gulandi_, Canarese; _Gulat-yelka_ of Wuddurs;
_Sora-panji-gadur_, Telegu of Yanadees; _Cofee-wattee-meeyo_,
Singhalese (this name seems to me a corruption of "coffee rat").
DESCRIPTION.--Fur thick and stiff, fulvous brown, mixed with black,
some olive brown mixed with fulvous, tawny grey beneath; hairs of
upper parts flattened, ashy grey, tipped yellow, with some thinner
and longer ones, also tipped yellow, with sub-terminal black band;
under fur soft and of a light lead colour; face and cheeks rough;
ears moderate, sub-ovate, hairy; tail round, tapering, scaly and
hairy, dark brown above, yellowish below; cutting teeth yellow.
SIZE.--Head and body, 4-1/2 inches; tail, 4 inches.
Dr. Kellaart says these are the rats most destructive to coffee-trees,
whole plantations being sometimes deprived of buds and blossoms by
them.
There is an illustration of one in Sir Emerson Tennent's 'Natural
History of Ceylon' in the act of cutting off the slender branches
which would not bear its weight in order to feed on the buds and
blossoms when fallen to the ground. "The twigs thus destroyed are
detached by as clean a cut as if severed with a knife." Sir Walter
Elliot writes of it: "The _gulandi_ lives entirely in the jungle,
choosing its habitation in a thick bush, among the thorny branches
of which, or on the ground, it constructs a nest of elastic stalks
and fibres of dry grass thickly interwoven. The nest is of a round
or oblong shape, from six to nine inches in diameter, within which
is a chamber about three or four inches in diameter, in which it rolls
itself up. Round and through the bush are sometimes observed small
beaten pathways along which the little animal seems habitually to
pass. Its motion is somewhat slow, and it does not appear to have
the same power of leaping or springing by which the rats in general
avoid danger. Its food seems to be vegetable, the only contents of
the stomach being the roots of the haryalee grass. Its habits are
solitary (except when the female is bringing up her young) and
diurnal, feeding in the mornings and evenings." Dr. Jerdon says:
"The Yanadees of Nellore catch this rat, surrounding the bush and
seizing it as it issues forth, which its comparatively slow actions
enable them to do easily. According to Sir Emerson Tennent the
Malabar coolies are so fond of their flesh that they evince a
preference for those districts in which the coffee-plantations are
subject to their incursions, where they fry the rats in cocoanut-oil
or convert them into curry." Both he and Dr. Kellaart mention the
migratory habits of this animal on the occurrence of a scarcity of
food. Kellaart says that in one day on such visits more than a
thousand have been killed on one estate alone.
NO. 379. GOLUNDA MELTADA.
_The Soft-furred Bush Rat_ (_Jerdon's No. 200_).
NATIVE NAMES.--_Mettade_, of Wuddurs; _Metta-yelka_, Telegu of
Yanadees; _Kera ilei_, Canarese.
HABITAT.--Southern India and Ceylon.
DESCRIPTION.--Fur very soft; above deep yellowish, olive brown or
reddish-brown, with a mixture of fawn; under fur lead colour; chin
and under parts whitish; head short; muzzle sharp; ears long and
hairy; tail shorter than body, scaly, but scales covered with short
black adpressed hairs; feet pale.
SIZE.--Head and body, 3-1/2 to 5-1/2 inches; tail, 2-1/4 to 4-1/4
inches.
The specific name of this rat is an absurd corruption, such as is
not unfrequent in Dr. Gray's names, of the native _mettade_, which
means soft. According to that accurate observer Sir Walter Elliot,
"the _mettade_ lives entirely in cultivated fields in pairs or small
societies of five or six;[25] making a very slight and rude hole in
the root of a bush, or merely harbouring among the heap of stones
thrown together in the fields, in the deserted burrow of the
_kok_,[26] or contenting itself with the deep cracks and fissures
formed in the black soil during the hot months. Great numbers perish
annually when these collapse and fill up at the commencement of the
rains. The monsoon of 1826 having been deficient in the usual fall
of rain at the commencement of the season, the _mettades_ bred in
such numbers as to become a perfect plague. They ate up the seed as
soon as sown, and continued their ravages when the grain approached
to maturity, climbing up the stalks of jowaree and cutting off the
ear to devour the grain with greater facility. I saw many whole fields
completely devastated, so much so as to prevent the farmers from
paying their rents. The ryots employed the Wuddurs to destroy them,
who killed them by thousands, receiving a measure of grain for so
many dozens, without perceptibly diminishing their numbers. Their
flesh is eaten by the Tank-diggers. The female produces six to eight
at a birth."--'_Madras Journ. Lit. Sc._' x. 1839.
[Footnote 25: In this case probably parents and young.]
[Footnote 26: _Nesokia providens_.]
Kellaart's _Golunda Newera_ is, I fancy, the same, although the
measurement he gives is less. Head and body, 3-1/4 inches; tail,
2-1/2. The description tallies, although Kellaart goes upon
difference in size and the omission of Gray to state that _G. meltada_
had the upper incisors grooved. He says that "this rat is found in
pairs in the black soil of Newara Elia, and is a great destroyer of
peas and potatoes." So its habits agree.
_GENUS HAPALOMYS_.
This was formed by Blyth on a specimen from Burmah of a murine animal
"with a long and delicately fine pelage and exceedingly long tail,
the terminal fourth of which is remarkably flattened and furnished
with hair more developed than in perhaps any other truly murine form;
limbs short, with the toes remarkably corrugated underneath; the
balls of the inguinal phalanges greatly developed, protruding beyond
the minute claws of the fore-feet, and equally with the more
developed claws of the hind-feet; head short; the ears small and
inconspicuous; the skull approaches in form that of _Mus
Indicus_,[27] but the rodential tusks are broader and flatter to the
front. Molars as in the _Muridae_ generally, but much worn in the
specimen under examination; they are considerably less directed
outward than usual, and the bony palate has therefore the appearance
of being narrow; the superorbital ridges project much outward in form
of a thin bony plate, and there is a considerable process at the base
of the zygoma anteriorly and posteriorly to the anti-orbital
foramen; zygomata broad, and compressed about the middle."
[Footnote 27: _Nesokia Blythiana_.]
NO. 380. HAPALOMYS LONGICAUDATUS.
HABITAT.--Shway Gheen, in the valley of the Sitang river in Burmah,
or its adjacent hills.
DESCRIPTION.--"Fur long and soft, measuring about five-eights of an
inch on the upper parts, slaty for the basal two-thirds, then
glistening brown with black tips, and a few long hairs of very fine
texture interspersed; lower parts dull white; whiskers black, long
and fine, and there is a tuft of fine blackish-hair anterior to the
ears."--_Blyth_.
SIZE.--Head and body of a male, 5-3/4 inches; tail 7-1/4 inches. Of
another specimen, female: 5-1/4 inches; tail, 7-1/2 inches; sole,
1-1/8 inch; ears posteriorly, 1-1/4 inch.
Specimens of adult male and female with a young one were forwarded
to the Asiatic Society's Museum by Major Berdmore.
* * * * *
We have now come to the end of the purely murine group as far as they
exist within the limits assigned to these investigations. I ought
perhaps to give some short notices of the following specimens
discovered in Thibet by the Abbe David, and described by Professor
Milne-Edwards in his 'Recherches sur les Mammiferes.'
NO. 381. MUS OUANG-THOMAE.
_The Kiangsi Rat_.
HABITAT.--Kiangsi in Thibet.
DESCRIPTION.--A tawny grey above, mixed with long hairs, tipped with
brown, greyish below; between the fore-paws a crescent of pure white,
which is a distinguishing mark of the species.
SIZE.--A little less than _Mus rattus_, which is about seven inches
long; tail an inch longer.
This rat Professor Milne-Edwards describes from a single specimen;
it is apparently rare, and was named after the Abbe David's Chinese
servant--'Recherches sur les Mammiferes,' p. 290.
NO. 382. MUS FLAVIPECTUS.
_The Yellow-breasted Rat_.
HABITAT.--Moupin; Thibet.
DESCRIPTION.--Reddish-brown; chin greyish; throat and chest tawny,
mixed with grey; belly and inside of limbs yellowish-grey; ears large,
nearly naked; incisors deep yellow; tail brown, covered with short
hairs.
SIZE.--About 7-3/4 inches; tail, 6-1/4 inches.--'Mammiferes,' p.
289.
NO. 383. MUS GRISEIPECTUS.
_The Grey-breasted Rat_.
HABITAT.--Moupin; Thibet.
DESCRIPTION.--Brown above; the under-parts of a clear grey.
SIZE.--About the same as the last, but with a somewhat shorter
tail.--'Mammiferes,' p. 290.
NO. 384. MUS CONFUCIANUS.
HABITAT.--Moupin; Thibet.
DESCRIPTION.--Fawn brown above, pure white below; lower part of
cheek white; on the back the fur is interspersed with longer hairs
of a blackish tint; feet pale.
SIZE.--Head and body, about 4 inches.--'Mammiferes,' p. 286.
NO. 385. MUS CHEVRIERI.
HABITAT.--Moupin; Thibet.
DESCRIPTION.--General colour tawny brown, grizzled with dark brown;
lower parts of a clear grey, almost white; ears short; feet small;
tail covered with short hair.
SIZE.--About 4-3/4 inches; tail about 3-1/2 inches.--'Mammiferes,'
p. 288.
NO. 386. MUS PYGMAEUS.
_The Pigmy Mouse_.
HABITAT.--Moupin; Thibet.
DESCRIPTION.--Distinguished by its very short ears and the square
form of its head; deep brown above; greyish-yellow beneath; tail
shorter than in the common mouse.
SIZE.--About 2-3/4 inches; tail, about 2 inches.--'Mammiferes,' p.
291.
ARVICOLINAE.
In this sub-family the molars are generally semi-rooted or rootless.
The _Arvicolinae_ or Voles consist of the American Musquash (_Fiber
zibethicus_), a very beaver-like water rat of large size; the
Lemmings (_Myodes_), of which there are several species which are
celebrated for their vast migrations; and the true Vole (_Arvicola_),
which is the only genus found in India, and then only in the colder
climate of the Himalayas. There are several species in Europe, of
which three are found in England. According to Professor Dallas, the
true Voles number about fifty species, arranged by various writers
under a considerable number of sub-genera. In India we have only
eight known species, and two more from the adjacent country of
Thibet.
[Illustration: Dentition of _Arvicola_.]
The European forms of _Arvicolae_ have been divided by Blasius into
four sub-genera of two divisions--the first division having rooted
molars in the adult animal--containing one sub-genus only,
_Hypudaeus_ of Illiger; the second division consists of three
sub-genera with rootless molars, viz. _Paludicola_, _Agricola_, and
_Arvicola_, which last has again been subdivided into long-eared and
short-eared Voles--_Arvicola_ and _Microtus_--distinguished by the
former having eight and the latter four mammae, and respectively six
and four tubercles on the plantae, the ears of the latter being almost
hidden by the fur.
None of the forms with which we have now to deal belong to the first
division, for, as far as the matter has been investigated, the Indian
Voles have rootless molars, but the character of the teeth in some
differs from the European forms, and therefore Mr. Blanford has
proposed a new section, _Alticola_, for their reception. I have not
space here, nor would it accord with the popular character of this
work, to go minutely into all the variation of dentition which
distinguish the different species. To those who wish to continue to
the minutest details the study of the Indian Voles, I recommend a
most careful and elaborate paper on them by Mr. W. T. Blanford, F.R.S.,
in the Journal of the Asiatic Society of Bengal, vol. L., pt. ii.;
but without entering into the microscopic particulars of each
species, I may here give a general idea of the formation of the teeth
of the _Arvicolae_ differing as it does so much from others of the
myomorphic or mouse-like group of rodents. In these the general
contour of the molar teeth is roundish oblong, the margins being wavy
or indented, according to the convolutions of the enamel, but in the
Voles there is a sharp angularity about these indentations; the
marginal lines, instead of being in well-rounded curves, are sharply
zigzag, forming acute angles. If you were to draw two close parallel
zigzag lines it would give you some idea of the contour of these teeth.
The molars are in fact composed of alternating triangular prisms,
with the outer folds of enamel forming deep and acute angles. The
other characteristics of this family are: skull, with brain case
rhomboidal, frontals much contracted; infra-orbital opening
typical; limbs moderate; tail moderate, or short and hairy.
_GENUS ARVICOLA_.
Muzzle blunt; fore-feet small, with short claws; soles naked; tail
longer than the hind-foot, clad with short hairs; incisors plain,
smooth in front. The fore-feet in some species have but a small wart
in place of a thumb; in others there is a small thumb with a minute
claw. The hind-feet have five toes.
NO. 387. ARVICOLA STOLICZKANUS.
_The Yarkand Vole_.
HABITAT.--Yarkand.
DESCRIPTION.--"Bright ferruginous brown above, pure white beneath;
fur soft, rather woolly, 0.5 to 0.6 inch long on the middle of the
back, the basal portion throughout both head and body being dark
leaden grey; this is the case on the back for about three-quarters
of the length of the hairs; the remaining quarter is rufous white,
tipped with darker rufous, whilst numerous rather longer hairs are
dark rufous-brown at the ends; rather a sharp line divides the rufous
of the back from the white belly; upper part of the head the same
colour as the back; upper whiskers dark brown, lower, including the
longest, white; ears small, rounded, hairy, completely concealed by
the fur, with rather short bright rufous hair near the margin inside;
and covered outside with longer and paler hair; feet small, the thumb
of the fore-foot quite rudimentary and clawless; remaining claws
long, compressed, sharply pointed, but much concealed by the long
white hairs which cover the upper part of the foot, sales naked;
tarsus hairy below, a few hairs between the pads of the toes; tail
short, apparently about a quarter the length of the body and head
together, covered with stiff fulvescent white hair, which extends
about half an inch beyond the end."--_W. T. Blanford_, 'Sc. Res. of
Second Yarkand Mission,' p. 43.
SIZE.--Head and body, about 4 inches; tail, with hair, 1-1/2.
NO. 388. ARVICOLA STRACHEYI.
_The Kumaon Vole_.
HABITAT.--Kumaon.
DESCRIPTION.--Light brown above, with a greyish tint and dusky
forehead; under-parts, feet, and tail white; ears small, not longer
than the fur, and thickly clad with hair; feet of moderate size; thumb
as in the last; tail short and covered with white hairs.
SIZE.--Head and body, about 3.7 inches; tail; 0.7.
This vole was procured first by Capt. (now Lieut.-Gen.) R. Strachey
at Kumaon.
NO. 389. ARVICOLA WYNNEI.
_The Murree Vole_.
NATIVE NAME.--_Kannees_.
HABITAT.--Northern Himalayas; Murree.
DESCRIPTION.--Dark brown above, with a slight greyish tinge; head
rufescent, and under-parts pale brown; tail dark brown; ears short
and rounded, hidden by the fur; fore-feet rather large; thumb small,
with a short claw; incisors orange.
SIZE.--Head and body, about 4-3/4 inches; tail 1-1/4 inch.
NO. 390. ARVICOLA ROYLEI.
_The Cashmere Vole_ (_Jerdon's No. 202_).
HABITAT.--Kashmir; Kunawur near Chini at 12,000 feet.
DESCRIPTION.--Yellowish-brown, with a rufous tint on the back, paler
below; tail brown above, whitish underneath; feet concolorous with
the under-part; ears small, hairy and nearly hidden by the fur;
incisors yellow in front.
SIZE.--Head and body, 3-3/4 inches; tail, 1-2/12 inch.
Jerdon states he got this vole at Kunawur, near Chini, again on the
south side of the Barendo pass, and also in the Pir Punjal.
NO. 391. ARVICOLA BLANFORDI.
_The Gilgit Vole_.
HABITAT.--Kashmir territory; Gilgit, at an elevation of 9000 to
10,000 feet.
DESCRIPTION.--Light greyish-brown above, slightly tinged with
rufous; greyish-white underneath; fur soft, the basal three-fourths
being slaty grey, the rest fawn colour, in some instances with black
tips, the hairs of the under-parts being white tipped; ears
moderately large, well above the fur, hairy; very long whiskers,
chiefly white, a few brown; feet whitish, moderate size; tail
cylindrical, not tapering, and well clad with hair, which project
about a fifth of an inch beyond the end of the vertebrae.
SIZE.--Head and body, about 4-1/2 inches; tail, 2 inches.
This vole was described by Dr. J. Scully in the 'Annals and Magazine
of Natural History,' for November, 1880, vol. vi., and he named it
after Mr. W. T. Blanford. It is said to be common on the mountains
around Gilgit.
* * * * *
The next two species come under the section _Paludicola_.
NO. 392. ARVICOLA BLYTHII.
HABITAT.--Western Thibet, Leh and Ladakh.
DESCRIPTION.--General colour above yellowish-brown, below pale
isabelline; fur soft; basal two-thirds of the upper hairs, and
one-half of the lower hairs, dark slaty; the upper hairs are tipped,
some isabelline and some, which are coarser and longer, dark brown;
ears round, small, equal, with the fur thinly clad with pale brown
hairs inside, and more thickly so with longer hairs outside; upper
whiskers dark brown, lower whitish; feet pale isabelline; soles
naked; tail cylindrical, distinctly ringed, covered with short light
brown hair like the under-parts in colour.
SIZE.--Head and body, about 3 to 4 inches; tail, 1 to 1-1/4 inch.
Mr. Blanford has written fully regarding this species, which was the
type of Blyth's genus _Phaiomys_, in the 'Scientific Results of the
Second Yarkand Mission,' page 39, in which he contends, after going
through a mass of literature on the subject, that there are no grounds
for constituting it the type of a new species; and, if this be
conceded, then the specific name given by Blyth, viz. _leucurus_,
being forestalled, it is necessary to rename it, which he has done
in honour of that well-known naturalist.
NO. 393. ARVICOLA MANDARINUS.
_The Afghan Vole_.
HABITAT.--Afghanistan; Chinese Mongolia.
DESCRIPTION.--Light greyish rufescent brown above, white beneath;
ears short, hidden by the fur and hairy; feet whitish; tail rufescent
brown.
SIZE.--About 4 inches; tail about 1 inch.
This vole, which is described and figured by Milne-Edwards, is
supposed to have been found in Afghanistan from a specimen in
Griffith's collection. _A. mandarinus_ comes from Chinese Mongolia,
and it is figured in the 'Recherches sur les Mammiferes.'
* * * * *
The next species was made a separate genus, _Neodon_, by Hodgson,
which has been adopted by Jerdon; but there are no good grounds for
continuing this separation. Mr. Blanford is certainly of this
opinion, and in his remarks on it (_see_ his 'Sc. Results Second
Yarkand Mission,' pp. 41-42) he writes: "The genus _Neodon_, appears
to be founded on characters of only specific importance, and the type
_N. Sikimensis_ is, I think, a true _Arvicola_."
NO. 394. ARVICOLA SIKIMENSIS.
_The Sikim Vole_ (_Jerdon's No. 203_).
NATIVE NAMES.--_Phalchua_, Nepalese, apparently Hindi; _Cheekyu_,
Kiranti; _Singphuci_, Thibetan.
HABITAT.--Nepal; Sikim; Thibet.
DESCRIPTION.--Fur soft and silky. "Deep brownish-black above with
a slight rusty shade, minutely and copiously grizzled with hairs of
a deep ferruginous tint" (_Horsfield_). Or a deep golden brown from
yellow hairs being intermixed; bluish-grey beneath, with a slight
fulvous tint; fur leaden grey for the basal three-fourths, the
terminal fourth being brownish or tawny with some tipped black; the
hairs of the under-parts are dipped with dirty white; ears project
beyond the fur moderately, and are hairy; feet very slender; tail
thinly clad with short brown hair. The female has six mammae.
SIZE.--Head and body, about 4-3/4 inches; tail, 1-1/2 inch.
Horsfield gives 5 inches for head and body.
According to Jerdon this vole has only been procured in Sikim near
Darjeeling, at heights varying from 7000 to 15,000 feet; but I
believe the area it inhabits to be much larger. Hodgson found his
specimens at Darjeeling, and on one occasion got a nest in a hollow
tree in the forest; it was saucer-shaped, of soft grass without any
lining, and contained a male, female, and two young. The latter were
"2-1/8 inches long, hairy above, nude below, and blind; the ears also
closed." Jerdon writes: "Mr. Atkinson found it under fallen trees
and stones on the top of Tonglo, near Darjeeling, 10,000 feet, whence
also I had a specimen brought me."
* * * * *
The next species is one described and figured by Professor
Milne-Edwards, and from Thibet he has two illustrations of it--one
of an entire blackish-brown, the other darker above, but with the
black belly.
NO. 395. ARVICOLA MELANOGASTER.
HABITAT.--Moupin in Tibet.
DESCRIPTION.--"It is characterised by the colour of the lower parts,
which are a blackish-grey. The upper parts are sometimes as black
as a mole, sometimes grizzled with brown" ('Mammiferes,' p. 284).
The brown specimen with the dark belly is evidently a rarity.
FAMILY SPALACIDAE.
The members of this family are characterised by very large incisors;
some have premolars, as in _Bathyergus_ and two other genera, but
not in the _Spalacinae_, of which our bamboo-rat (_Rhizomys_) is the
representative in India. "The grinding teeth are rooted, not
tuberculate, but with re-entering enamel folds; infra-orbital
opening moderate or small, with no perpendicular plate; occipital
plane high, often sloped boldly forward; palate narrow; form
cylindrical; eye and ear-conch very small, sometimes rudimentary;
limbs short and stout; claws large; tail short or absent" (_Alston_,
'P. Z. S.' 1876, p. 86). There are two subfamilies--_Spalacinae_ and
_Bathyerginae_.
_GENUS RHIZOMYS--THE BAMBOO-RAT_.
"Form robust; eyes very small; ears very short, naked; pollex
rudimentary; tail rather short, partially haired; skull broad;
occipital plane only slightly sloped forward; infra-orbital opening
small, sub-triangular; upper incisors arched forward; no premolar;
upper molars with one deep internal and two or more external
enamel-folds; the lower molars reversed."--_Alston_.
NO. 396. RHIZOMYS BADIUS.
_The Chestnut Bamboo-Rat_ (_Jerdon's No. 201_).
NATIVE NAME.--Known to the Chingpaws or Kakhyens as the
_Yewcron_.--_Anderson_.
HABITAT.--The Sikim and Nepal Terai; Burmah; Arakan; Kakhyen Hills.
[Illustration: _Rhizomys badius_.]
DESCRIPTION.--Fine fur, of a grey or slaty grey for two-thirds of
the basal portion, the remaining upper third being from a deep to
a bright chestnut. "Most intense on the head, and dullest on the rump"
(_Anderson_). "Below dark ashy grey" (_Jerdon_). "The fur of the
under-parts in these Eastern examples of the species" (referring to
those from the Kakhyen hills) "is paler and more reddish than
chestnut, whereas in some Nepal animals it inclines even to slaty
grey, washed with reddish. The area immediately around the muzzle
and the chin is pale brownish, with a tinge of greyish, and the teeth
are brilliant reddish, the nose, ears, feet, and tail being pale
flesh-coloured" (_Anderson_, 'Anat. and Zool. Res.' p. 329).
SIZE.--Head and body, 7 inches; tail, about 2-1/2 inches.
Jerdon says of this species that "it eats the roots of bamboos and
other trees, constructing burrows under the roots. It is said to be
very bold, and easily taken." "In Burmah it constructs its burrows
amongst a rank and tall jungle grass, on the roots of which it is
said to live" (_Anderson_). Blyth, who writes of the Burmese form,
says: "it is barely separable from _R. badius_, from which it seems
to differ only in its much brighter colouring."
NO. 397. RHIZOMYS ERYTHROGENYS.
_The Red-cheeked Bamboo-Rat_.
HABITAT.--Burmah; the Salween hill tracts; Tenasserim.
DESCRIPTION.--Upper parts dark iron grey; almost black on the top
of the head; the upper lip, chin and upper part of the throat are
white, also the chest and belly, which are however more or less tinged
with grey and reddish; the lower portion of the throat is dark grey;
the sides of the head and cheeks are bright golden red; the feet are
sparsely clad and leaden coloured, except the toes of the hind feet,
which are fleshy white; tail rather thick at the base, quite naked,
not scaly, and of a leaden hue; claws rather broad, and moderately
strong.
SIZE (of the living female).--Head and body, 14-3/4 inches; tail,
5.35 inches.
Dr. Anderson, from whose work I have taken the above description,
and who was the first to describe and name this animal, says that
a female was recently received in the Zoological Gardens from Mr.
A. H. Hildebrand.
NO. 398. RHIZOMYS PRUINOSUS.
_The Hoary Bamboo-Rat_.
HABITAT.--Assam; very common about Cherrapoonjee; Burmah; Kakhyen
hills east of Bhamo.
DESCRIPTION.--Brown above, grizzled with white; the base of the fur
being slaty grey, tipped with brown, and intermixed with longer hairs,
terminating in white bands; underneath much the same, only the
white-tipped hairs are shorter and less numerous; whiskers dark
brown; the head is generally more grey; ears, nose, feet and tail
of a dusky flesh tint; tail one-third of the body.
SIZE.--Head and-body, about 11 to 13 inches; tail, 3 to 4 inches.
NO. 399. RHIZOMYS MINOR.
_The Small Bamboo-Rat_.
NATIVE NAME.--_Khai_, Aracanese.
HABITAT.--Burmah, Upper Martaban, and at Yanageen on the
Irrawaddy.--_Blyth_.
DESCRIPTION.--"Dark sooty brown above, slightly tinged with deep
umber, which is most distinct on the sides of the head and neck, and
in reflected light; the under parts are like the upper, only the brown
tint is almost absent; the whiskers are black, and tail very sparsely
haired" (_Anderson_). "Dusky brown colour, with white muzzle and
around the eye, and pale naked feet" (_Blyth_).
SIZE.--Head and body, 6-1/2 inches; tail, 1-3/4 inch.
Blyth says he obtained a living specimen in Upper Martaban, and
recognised it as the same as what had been obtained in Siam. The Rev.
Mr. Mason writes of it: "This animal, which burrows under old bamboo
roots, resembles a marmot more than a rat; yet it has much of the
rat in its habits. I one night caught a specimen gnawing a cocoa-nut,
while camping out in the jungles."
* * * * *
I may here mention a curious little animal, which is apparently a
link between the MURIDAE and the SPALACIDAE, _Myospalax
fuscocapillus_, named and described by Blyth ('J. A. S. B.' xv. p.
141), found at Quetta, where it is called the "Quetta mole." A full
account of it by Mr. W. T. Blanford is to be found in the 'Journal
Asiatic Society of Bengal,' (vol. L. pt. ii.).
FAMILY DIPODIDAE.
This family contains a form of rodent similar to, yet more pronounced
than, the jerboa rats, of which I have already treated. It includes
the true Jerboas (_Dipus_), the American Jumping Mice (_Zapus_), the
_Alactaga_, and the Cape Jumping Hare (_Pedetes caffer_). The
characteristics of the family are as follows:--
"Incisors compressed; premolars present or absent; grinding teeth
rooted or rootless, not tuberculate, with more or fewer transverse
enamel folds; skull with the brain-case short and broad;
infra-orbital opening rounded, very large (often as large as
the orbit); zygomatic arch slender, curved downwards; the malar
ascending in front to the lachrymal in a flattened perpendicular
plate; facial surface of maxillaries minutely perforated; mastoid
portion of auditory bullae usually greatly developed; metatarsal
bones elongated, often fused into a cannon bone; form gracile; front
portion of body and fore-limbs very small; hind limbs long and strong,
with from three to five digits; tail long, hairy. Three sub-families"
(_Alston_ On the Order GLIRES, 'P. Z. S.' 1876). The three
sub-families are _Zapodidae_,[28] _Dipodinae_ and _Pedetinae_, but
we have only to deal with the second.
[Footnote 28: Formerly _Jaculinae_.]
[Illustration: Dentition of Jerboa.]
_GENUS DIPUS--THE JERBOAS_.
Hind feet with three digits; tail cylindrical and tufted; incisors
grooved; premolars absent, or, if found, then in the upper jaw and
rudimentary; skull with very broad occipital region; greatly
developed auditory bullae; the cervical vertebrae are more or less
anchylosed, and the metatarsals are united. They are not found in
the plains of India, though one species inhabits Yarkand, and two
more are found in Eastern Persia.
[Illustration: _DIPUS_.]
NO. 400. DIPUS LAGOPUS.
_The Yarkand Jerboa_.
HABITAT.--Koshtak, south of Yarkand; Yarkand; and
Yangihissar.--_Blanford_.
DESCRIPTION.--"Colour above light sandy brown, slightly washed with
dusky, below pure white; a white band across the outside of the thigh;
tail pale brown above, whitish below, with a tuft of longer hair,
altogether about 2-1/2 inches long; at the end the terminal portion
pure white, the proximal portion black or dark-brown on the upper
part and sides, but brown or white beneath the tail. The fur is very
soft and rather long, 0.6 to 0.8 inch in the middle of the back; on
the upper parts it is ashy grey at the base and for the greater parts
of its length, pale sandy brown near the end; the extreme tip dusky
brown; on the lower parts it is white throughout; ears about half
the length of the head, oval, naked inside, thinly clothed with short
brown hair outside; face sandy; the hairs grey at the base; sides
of head whitish; whiskers as usual very long, exceeding three inches;
the uppermost brown; the longest white, except at the base; the lower
entirely white; the long hairs beneath the hind feet all white, as
are the feet throughout."--_Blanford_, 'Sc. Res. of Sec. Yarkand
Mission,' pp. 58,59.
_GENUS ALACTAGA_.
"Hind feet with _five_ digits, of which the first and fifth do not
reach the ground; tail cylindrical, tufted; skull with the occipital
region less broad, and the auditory bullae smaller; infra-orbital
opening with no separate canal for the nerve; incisors plain. One
very small premolar present above only."--_Alston_.
NO. 401. ALACTAGA INDICA.
NATIVE NAME.--_Khanee_, Afghan.
HABITAT.--Afghanistan; Eastern Persia.
DESCRIPTION.--Fawn colour above; the hair with black tips and ashy
grey at the base; under-parts white; upper parts of thigh white; a
black spot behind and inside the thigh just below the white;
remainder of the outside and lower part of the inside of the thighs
brown; a white line running down the front, and extending over the
upper portion of the tarsi and feet; proximal portion of tarsus brown
at the sides. (_See_ 'Blanford's Eastern Persia,' vol. ii. p. 77.)
The tail is brown with a white tip; ears thinly clad with brown hairs;
head brown above, whitish around the eyes; whiskers black.
SIZE.--Head and body, 3-1/2 inches; tail, 7 inches.
This animal is unfortunately named, as it is not Indian at all;
equally unfortunate, as Mr. Blanford has shown, is Blyth's name
_Bactrianus_, for it does not inhabit that tract, so the original
title stands. Hutton, in his 'Rough Notes on the Zoology of Candahar'
('J. A. S. B.' xv. p. 137), writes of it as follows: "This beautiful
little animal is abundant over all the stony plains throughout the
country, burrowing deeply, and when unearthed bounding away with
most surprising agility after the manner of the kangaroo-rat. It is
easily tamed, and lives happily enough in confinement if furnished
with plenty of room to leap about. It sleeps all day, and so soundly
that it may be taken from its cage and examined without awaking it;
or at most it will half open one eye in a drowsy manner for an instant,
and immediately close it again in sleep. It retires to its burrows
about the end of October, and remains dormant till the following
April, when it throws off its lethargy and again comes forth." There
is a good engraving of this animal in Cassel's new Natural History.
* * * * *
We have now closed our account of the Myomorpha or Mouse-like Rodents,
and will proceed to the next Section, HYSTRICOMORPHA, or
Porcupine-like Rodents.