THE GENUS CLUBIONA

These spiders are all pale and most of them without markings. The eyes

are close to the front edge of the head and cover more than half its

width (figs. 50, 54, 56). The upper row is longer and the eyes larger

and the middle pair farthest apart. The distance between this pair

varies according to the species. In _crassipalpis_ and _canadensis_ it

is little more than that between the middle and lateral eyes, while in

_rubra_ it is nearly twice as great. The mandibles of the females are

swelled at the base in front, and this swelling is greatest in

_canadensis_ (fig. 55). The mandibles of the males are longer and are

shaped in a variety of ways according to the species. The shape of the

epigynum is indistinct and variable, and females of different species

are difficult to distinguish. The females of _ornata_ and _excepta_ are

known by their markings and those of _rubra_ by their size and

resemblance to the male. The females of _crassipalpis_ and _tibialis_

are doubtful. The palpal organs and male palpi are of great variety and

distinguish the males of all species without much difficulty.

[Illustration: FIG. 48. Female Clubiona crassipalpis, enlarged four

times.]

The Clubionas live in flat tubes of silk on leaves of low plants in

summer and under bark and stones at all seasons.

$Clubiona crassipalpis.$--A quarter of an inch long and pale, without

markings. The head is sometimes a little darker than the rest of the

body, and the mandibles and ends of the male palpi are always darker.

The eyes of the upper row are almost equidistant, the middle pair only a

little farther apart than they are from the lateral eyes. The mandibles

of the male (figs. 49, 50) are elongated as usual, narrowed toward the

end, and thickened in front just above the middle. On the outer side in

front is a sharp ridge that extends from the base of the claw halfway up

the mandible. The inner edges of the mandibles are thin and inclined

backward toward the mouth, but there is no line or ridge between the

thick and thin portions as in some other species. The palpi of the male

(fig. 51) have the patella and tibia both short. The tibia is widened on

the outer side and laps over the tarsus, extending in a blunt hook for

half its length.

[Illustration: FIGS. 49, 50, 51, 52. Clubiona crassipalpis.--49, head

and mandibles of male from the left side. 50, head and mandibles of male

from in front. 51, palpus of male. 52, maxillæ, labium, and ends of

mandibles.]

[Illustration: FIGS. 53, 54. Clubiona tibialis.--53, end of palpus of

male, showing large tibia. 54, head and mandibles of male.]

$Clubiona tibialis.$--Quarter of an inch long, the same size and color

as _crassipalpis_, with no markings and no dark color except on the

mandibles and male palpi. The middle eyes are a little farther apart

than in _crassipalpis_. The male mandibles (fig. 54) are narrower than

in _crassipalpis_ and without the sharp ridge on the outer side, but on

the inside they are sharply hollowed out with a ridge between the thick

outer and thin inner portions. The male palpi (fig. 53) have the patella

about as long as wide, as it is in _crassipalpis_ and _canadensis_, but

the tibia is very large, with a hook in the usual place on the outer

side and a much larger process, which extends forward over the tarsus.

The tarsus is long and thickened in the middle where it rests against

the process of the tibia.

[Illustration: FIGS. 55, 56, 57. Clubiona canadensis.--55, head of

female, showing the swelled mandibles. 56, head and mandibles of male.

57, palpus of male.]

$Clubiona canadensis.$--Quarter of an inch long and without markings.

The upper eyes are nearly equidistant, as they are in _crassipalpis_,

but the legs and palpi are shorter. The male mandibles (fig. 56) are

much like those of _crassipalpis_, but have not so sharp a ridge on the

outer side. The male palpi (fig. 57) have the patella longer than wide

and the tibia wider than long, with two processes on the outer side. The

upper tibial process is a simple point extending along the outer side of

the tarsus for a quarter of its length. The under process is twice as

long, with a projecting corner at its base and running forward to a

sharp point, with a round notch halfway between the point and base. The

female has the head wider and the mandibles very much swelled in front

at the base (fig. 55). The fourth leg is longest and about as long as

the body.

[Illustration: FIGS. 58, 59. Clubiona rubra.--58, front of head and

mandibles of male. 59, palpus of male.]

$Clubiona rubra.$--Smaller than the other species, a sixth to a fifth of

an inch long. Cephalothorax light yellow brown, darker toward the front.

Abdomen pale in front and darkened with brownish red at the sides and

behind and along the middle of the back. The hind middle eyes are nearly

twice as far apart as they are from the lateral eyes. The fourth legs

are longest in both sexes, and there is little difference in the length

of the legs of the two sexes. The mandibles of the female are but little

swelled in front, not much more than those of the male, and the male

mandibles (fig. 58) are only a little narrowed at the ends. The male

palpi (fig. 59) have the tibia very much widened on the outer side, with

a short tooth in the middle and two larger short processes on the outer

side. The palpal organ has a large dark-colored process in the middle.

The epigynum is pointed behind, with a notch in the middle and two black

spots under the skin toward the front of the abdomen.

[Illustration: FIGS. 60, 61, 62. Clubiona ornata.--60, back of female

enlarged four times to show markings. 61, palpus of male. 62, front of

head and mandibles of male.]

$Clubiona ornata.$--This is one of the few Clubionas that have markings

on the abdomen. The general color is pale as usual, but the abdomen has

a dark stripe in the middle, broken into spots behind and bordered by

pale yellow. At the sides are oblique dark and light transverse

markings. These marks are of different sizes in different individuals

and connected in different ways. The length is from a third to half an

inch. The abdomen is wide across the middle and more pointed behind than

in most species. The front middle eyes are about as far apart as in

_tibialis_ and nearer than in _rubra_. The mandibles are not much

swelled in front. The legs of the male are longer than those of the

female, with the first pair longest, while in the female the fourth pair

is longest. The male mandibles (fig. 62) are narrow at the end and

hollowed on the inner edges as in _tibialis_, with a sharp ridge between

the thick and the thin portions. The male palpi (fig. 61) have two

processes on the outer side of the tibia longer than in _rubra_ and

shorter than in _canadensis_. The tibia is a little widened toward the

end and curved outward.

[Illustration: FIGS. 63, 64, 65. Clubiona excepta.--63, front of head

and mandibles of male. 64, palpus of male. 65, back of female enlarged

four times to show markings.]

$Clubiona excepta.$--A third of an inch long and with very distinct gray

markings on the abdomen. The cephalothorax is light yellow brown, and

the legs are more deeply colored with yellow than in most species. The

abdomen is white, with sometimes a yellow mark on the front of the

abdomen, and on the hinder half of the abdomen is a middle row of gray

spots and a row of larger transverse spots on each side. The fourth legs

are longest in both sexes. The male palpi have the tibia longer than

usual and patella and tibia about the same length. The tibia has a small

process with two teeth (fig. 64). The tarsus is oval and narrow and the

palpal organ small.

[Illustration: FIG. 66. Trachelas ruber, enlarged four times.]

$Trachelas ruber.$--One-third of an inch long, with the cephalothorax

short and wide like Clubiona, and the deep orange-brown color of

Dysdera. The cephalothorax is almost as wide as long, and widest

opposite the second legs. The head is three-quarters as wide as the

thorax and as high in the middle halfway between the eyes and the dorsal

groove. The front of the head is low, as in Clubiona. The front row of

eyes is a little curved, so that the middle pair are half their diameter

higher than the lateral. The upper row is much longer and the lateral

eyes are farther from the middle than these are from each other. The

labium and maxillæ are like those of Clubiona and the palpi very

slender. The first pair of legs are thicker than the others and as long

as the fourth. The second pair are also somewhat thickened. The legs are

orange brown, darkest on the front pair. The cephalothorax is dark brown

and finely roughened over the whole surface, without hairs except in

front. The abdomen is pale, with no markings except over the dorsal

vessel and the muscular spots. Some light-colored individuals have all

the colors paler. Under stones and leaves.

[Illustration: FIGS. 67, 68, 69. Chiracanthium viride.--67, female

enlarged four times. 68, eyes from in front. 69, maxillæ, labium, and

mandibles from below.]

$Chiracanthium viride.$--This has the color and general appearance of

the Clubionas, but the legs are longer and the first legs are

considerably longer than the fourth. The body is shorter and the abdomen

is wider and thicker in the middle. The female (fig. 67) is a third of

an inch long and the front legs two-fifths of an inch. The eyes (fig.

68) are arranged as in Clubiona. The maxillæ and labium are like those

of Clubiona, but the sternum is shorter and rounder. The head is but

little narrowed and the eyes cover almost its whole width. The upper

spinnerets are longer than the lower and distinctly two-jointed. The

spines of the legs are small and inconspicuous. The color in life is

greenish white, the mandibles brown, and the stripe over the dorsal

vessel darker than the rest of the abdomen.

The male has the front legs nearly three times as long as the body,

though the other legs are not much longer than in the female. The

mandibles are also elongated, as in the males of Clubiona. The male

palpi have the tarsus long, with a pointed process that extends backward

over the tibia between two processes on that joint.

The Common Spiders of the United States · 第 20 / 46 题